Heyrman, J; Verbeeren, J; Schumann, P; Swings, J; De Vos, P (July 2005). "Six novel Arthrobacter species isolated from deteriorated mural paintings". International Journal of Systematic and Evolutionary Microbiology. 55 (Pt 4): 1457–64. doi:10.1099/ijs.0.63358-0. PMID 16014466. https://doi.org/10.1099%2Fijs.0.63358-0