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Two Medicine Formation
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<h2 id="history-of-research">History of research</h2> <p>In <a href="/facts/1913_in_paleontology/3DxrWEaN">1913</a>, a <a href="/facts/US_Geological_Survey/nVaGtMjl">US Geological Survey</a> crew headed by Eugene Stebinger and a <a href="/facts/US_National_Museum/nn6mUbfE">US National Museum</a> crew headed by <a href="/facts/Charles_W._Gilmore/Qt8GVChj">Charles Gilmore</a> worked together to excavate the first dinosaur of the formation.<a class="footnote-ref" id="fnref:1" href="#fn:1"><sup>1</sup></a> Stebinger was the first to identify the Two Medicine Formation and formally described the first fossils in a scientific paper published in <a href="/facts/1914_in_paleontology/2woKqXj8">1914</a>.<a class="footnote-ref" id="fnref:2" href="#fn:2"><sup>2</sup></a> Gilmore returned to the Formation in <a href="/facts/1928_in_paleontology/Kas5A1ex">1928</a> and <a href="/facts/1935_in_paleontology/g0JKVp0r">1935</a>.<a class="footnote-ref" id="fnref:3" href="#fn:3"><sup>3</sup></a> During this time frame only three species were named and of these only <i><a href="/facts/Styracosaurus_ovatus/UuQQoD6I">Styracosaurus ovatus</a></i> and <i><a href="/facts/Edmontonia_rugosidens/1222bGhF">Edmontonia rugosidens</a></i> are still regarded as valid.<a class="footnote-ref" id="fnref:4" href="#fn:4"><sup>4</sup></a> <a href="/facts/Barnum_Brown/aI9FmlhV">Barnum Brown</a> prospected the formation in <a href="/facts/1933_in_paleontology/phKmFdsa">1933</a>, but found nothing significant.<a class="footnote-ref" id="fnref:5" href="#fn:5"><sup>5</sup></a> Both of their research were interrupted by <a href="/facts/World_War_II/2efV5L1U">World War II</a>.<a class="footnote-ref" id="fnref:6" href="#fn:6"><sup>6</sup></a> In <a href="/facts/1977_in_paleontology/SPR8VKkN">1977</a>, Trexler reports finding <a href="/facts/Hadrosaur/v4t7JSPB">hadrosaur</a> remains west of <a href="/facts/Choteau%2c_Montana/d1HYSBK9">Choteau, Montana</a>.<a class="footnote-ref" id="fnref:7" href="#fn:7"><sup>7</sup></a> During the <a href="/facts/1978_in_paleontology/qW7wYXAM">next year</a> baby hadrosaurs were discovered.<a class="footnote-ref" id="fnref:8" href="#fn:8"><sup>8</sup></a> In <a href="/facts/1979_in_paleontology/W3WSLGoS">1979</a>, Horner and Makela referred these hadrosaur bones to <i><a href="/facts/Maiasaura_peeblesorum/6eqMzunc">Maiasaura peeblesorum</a></i>.<a class="footnote-ref" id="fnref:9" href="#fn:9"><sup>9</sup></a> The announcement attracted renewed scientific interest to the formation and many new kinds of dinosaurs were discovered.<a class="footnote-ref" id="fnref:10" href="#fn:10"><sup>10</sup></a> More nesting sites were discovered later, including the Devil's Coulee site yielding <i><a href="/facts/Hypacrosaurus_stebingeri/BeeBEEva">Hypacrosaurus stebingeri</a></i> in southern <a href="/facts/Alberta/PjKJFbJY">Alberta</a> in <a href="/facts/1987_in_paleontology/HqEKF1EX">1987</a>. </p> <h2 id="geology">Geology</h2> <p>The loosely consolidated fine grain sediments composing the formation allow for fast plant growth in <a href="/facts/Badlands/14sa2rpz">badland</a> areas, limiting the number of exposed outcrops.<a class="footnote-ref" id="fnref:11" href="#fn:11"><sup>11</sup></a> <a href="/facts/Paleosols/K5jzLBxf">Paleosols</a>, <a href="/facts/Fluvial/nkLrSheo">fluvial</a> deposits and <a href="/facts/Bentonitic/N9Hb8lX9">bentonitic</a> layers are common in the Two Medicine Formation.<a class="footnote-ref" id="fnref:12" href="#fn:12"><sup>12</sup></a> </p> <h3>Age</h3> <p>The Two Medicine Formation spans from 82.4 to 74.4 Ma, nearly the entire length of the <a href="/facts/Campanian/MoPSxcqL">Campanian</a> stage. The age of the different members of the formation are as follows:<a class="footnote-ref" id="fnref:13" href="#fn:13"><sup>13</sup></a> </p> <ul><li>Rock City Member: 82.4-80.8 Ma</li> <li>Shields Crossing Member: 80.8-80.2 Ma</li> <li>Hagans Crossing Member: 80.2-77 Ma</li> <li>Flag Butte Member: 77-74.8 Ma</li></ul> <h3>Equivalents</h3> <p>There are several equivalents to the Two Medicine Formation, as with many geologic formations (most of which are named after their <a href="/facts/Type_locality_(geology)/ky6QjASt">type locality</a>). The Sweetgrass Arch in Montana divides the Two Medicine from the <a href="/facts/Judith_River_Formation/mbTI3gLp">Judith River Formation</a>, <a href="/facts/Bearpaw_Shale/ycaYRfIq">Bearpaw Shale</a>, <a href="/facts/Claggett_Shale/SVWusy6H">Claggett Shale</a>, and <a href="/facts/Eagle_Sandstone/Lg0IHDLR">Eagle Sandstone</a>. Across the <a href="/facts/Canada%25E2%2580%2593United_States_border/Q7TTKItn">Canada–United States border</a>, the Two Medicine Formation correlates to the <a href="/facts/Belly_River_Group/8mYQvyHD">Belly River Group</a> in southwest Alberta, and the <a href="/facts/Pakowki_Formation/F0dXUW7p">Pakowki Formation</a> eastward. </p> <h3>Stratigraphy</h3> <p>The Two Medicine overlies the <a href="/facts/Virgelle_Sandstone/HVrzxKr1">Virgelle Sandstone</a>, which formed from the beach sands exposed on northern and western shores of the receding Colorado Sea.<a class="footnote-ref" id="fnref:14" href="#fn:14"><sup>14</sup></a> A Cretaceous <a href="/facts/Interior_Seaway/I4AKChV9">Interior Seaway</a> transgression submerged the area briefly early on in Two Medicine history leaving anomalous paralic sediments and isolated <a href="/facts/Shale/bsIhqY5F">shale</a> bodies about 100 m above the base of the formation.<a class="footnote-ref" id="fnref:15" href="#fn:15"><sup>15</sup></a> The Middle portion of the two medicine formation is about 225 m thick, deposited while the Clagette Sea was receding and the <a href="/facts/Bearpaw_Sea/I4AKChV9">Bearpaw Sea</a> transgressing.<a class="footnote-ref" id="fnref:16" href="#fn:16"><sup>16</sup></a> This portion is stratigraphically equivalent to the <a href="/facts/Judith_River_Formation/mbTI3gLp">Judith River Formation</a> and <a href="/facts/Judith_River_Group/JKT3RG6c">Judith River Group</a>.<a class="footnote-ref" id="fnref:17" href="#fn:17"><sup>17</sup></a> The sediments are mainly <a href="/facts/Bentonitic/N9Hb8lX9">bentonitic</a> <a href="/facts/Siltstones/sI119qT3">siltstones</a> and mudstones with "occasional <a href="/facts/Sandstone/RP8B6fqL">sandstone</a> lenses."<a class="footnote-ref" id="fnref:18" href="#fn:18"><sup>18</sup></a> These sediments are thought to be the remains of a coastal plain "far removed" from the interior sea.<a class="footnote-ref" id="fnref:19" href="#fn:19"><sup>19</sup></a> The upper portion is about one half of the formation.<a class="footnote-ref" id="fnref:20" href="#fn:20"><sup>20</sup></a> Its sediments are similar to the middle portions but punctuated by extensive red beds and <a href="/facts/Caliche/MATKYcUo">caliche</a> horizons.<a class="footnote-ref" id="fnref:21" href="#fn:21"><sup>21</sup></a> The uppermost 80 m were deposited after the inundation of the Judith River equivalent sediments by the Bearpaw Sea.<a class="footnote-ref" id="fnref:22" href="#fn:22"><sup>22</sup></a> They are thought to have been deposited in only 500,000 years.<a class="footnote-ref" id="fnref:23" href="#fn:23"><sup>23</sup></a> Bentonitic ash is common in the Two Medicine.<a class="footnote-ref" id="fnref:24" href="#fn:24"><sup>24</sup></a> To the south extrusive volcanic activity occurred in association with the <a href="/facts/Boulder_Batholith/neDRJsda">Boulder Batholith</a> collectively called the <a href="/facts/Elkhorn_Mountains/gljFoUX0">Elkhorn Volcanics</a>.<a class="footnote-ref" id="fnref:25" href="#fn:25"><sup>25</sup></a> </p> <h3>Taphonomy</h3> <p>Most of the vertebrate fossils are preserved by <a href="/facts/CaCo3/UQ1wLFna">CaCo3</a> <a href="/facts/Permineralization/IquXFKhR">permineralization</a>.<a class="footnote-ref" id="fnref:26" href="#fn:26"><sup>26</sup></a> This type of preservation preserves high levels of detail, even down to the microscopic level.<a class="footnote-ref" id="fnref:27" href="#fn:27"><sup>27</sup></a> However, it also leaves specimens vulnerable to <a href="/facts/Weathering/uSYGlv0H">weathering</a> when exposed to the surface.<a class="footnote-ref" id="fnref:28" href="#fn:28"><sup>28</sup></a> </p> <h2 id="paleoenvironment">Paleoenvironment</h2> <h3>Climate</h3> <p>The Two Medicine Formation was deposited in a seasonal, semi-arid climate with possible <a href="/facts/Rain_shadow/aRWcxnWo">rainshadows</a> from the Cordilleran highlands. This region during the Campanian experienced a long dry season and warm temperatures. Lithologies, <a href="/facts/Invertebrate/3CyqCVnN">invertebrate</a> faunas, and plant and pollen data support the above interpretation. The extensive red beds and <a href="/facts/Caliche/MATKYcUo">caliche</a> horizons of the upper Two Medicine are evidence of at least seasonally arid conditions.<a class="footnote-ref" id="fnref:29" href="#fn:29"><sup>29</sup></a> Some of the dinosaurs from the formation have been speculated to have shown signs of drought-related death.<a class="footnote-ref" id="fnref:30" href="#fn:30"><sup>30</sup></a> </p> <h3>Elevation</h3> <p>A more upland environment existed in the south of the Two Medicine Formation.<a class="footnote-ref" id="fnref:31" href="#fn:31"><sup>31</sup></a> Streams had a northeasterly flow away from these southwestern uplands.<a class="footnote-ref" id="fnref:32" href="#fn:32"><sup>32</sup></a> The southern part of the Two medicine formation grades into brackish water siltstone/sandstone series called the <a href="/facts/Horsethief_Formation/qhaUEaPd">Horsethief Formation</a>.<a class="footnote-ref" id="fnref:33" href="#fn:33"><sup>33</sup></a> The sediments of the Horsethief represent shallower water deposits than the Bearpaw Shale adding further evidence of higher elevation areas existing in the south.<a class="footnote-ref" id="fnref:34" href="#fn:34"><sup>34</sup></a> </p> <h2 id="egg-mountain-site">Egg Mountain site</h2> <p>Egg Mountain, which is near <a href="/facts/Choteau%2c_Montana/d1HYSBK9">Choteau, Montana</a>, was discovered in 1977 by Marion Brandvold, owner of the Trex Agate Rock Shop in <a href="/facts/Bynum%2c_Montana/qeEpXEyD">Bynum, Montana</a>, who discovered the bones of juvenile dinosaurs at this site. It is a colonial nesting site on the <a href="/facts/Willow_Creek_(Montana)/L2p4yhVW">Willow Creek</a> <a href="/facts/Anticline/FQDWhobn">Anticline</a> in the Two Medicine Formation that is famous for its fossil eggs of <a href="/facts/Maiasaura/6eqMzunc">Maiasaura</a>, which demonstrated for the first time that at least some dinosaurs cared for their young. The eggs were arranged in dug-out earthen nests, each nest about a parent's body length from the next, and baby dinosaurs were also found with skeletons too cartilaginous for them to walk - similar to those of <a href="/facts/Altricial/C8JqqyaX">altricial</a> (helpless) baby <a href="/facts/Birds/RJF6zwLP">birds</a>. The parent(s) must then have brought food to the young, and there is plant matter in the nests that may be evidence of either this or for incubation of the eggs. <i>Maiasaura</i> also grew extremely fast, at rates comparable to modern birds. Skeletons of <i><a href="/facts/Orodromeus/hJ7tFcFI">Orodromeus</a></i> and skeletons and eggs of <i><a href="/facts/Troodon/cRJams94">Troodon</a></i> were also found at Egg Mountain. </p> <h2 id="biostratigraphy">Biostratigraphy</h2> <p>Most <a href="/facts/Dinosaur-bearing_rock_formations/G8KXXsw0">dinosaur-bearing rock formations</a> do not contain multiple distinct faunas at different positions within the formation's stratigraphic column.<a class="footnote-ref" id="fnref:35" href="#fn:35"><sup>35</sup></a> Usually the lower sediments of a given formation will contain the same kinds of dinosaurs as the upper sediments, or the species composition changes only gradually.<a class="footnote-ref" id="fnref:36" href="#fn:36"><sup>36</sup></a> However, some researchers had argued that the Two Medicine Formation was an exception, preserving multiple distinct dinosaur faunas.<a class="footnote-ref" id="fnref:37" href="#fn:37"><sup>37</sup></a> </p><p>Later research came to find that the supposedly distinct dinosaur faunas at different levels of the formations were more similar than had been previously thought.<a class="footnote-ref" id="fnref:38" href="#fn:38"><sup>38</sup></a> While the dinosaur fauna of the lower and middle sections Two Medicine was apparently diverse, the quality of preservation was low and few of these remains can be referred to individual species.<a class="footnote-ref" id="fnref:39" href="#fn:39"><sup>39</sup></a> The middle Two Medicine is a better source of fossils, but still poor overall. This makes it difficult to argue that these sections of the formation preserve distinct faunas.<a class="footnote-ref" id="fnref:40" href="#fn:40"><sup>40</sup></a> </p><p>The upper portion of the formation is more diverse and preserves better quality fossils.<a class="footnote-ref" id="fnref:41" href="#fn:41"><sup>41</sup></a> However, many of the taxa that supposedly distinguished it as a separate fauna have since been found in older sediments. In particular, <i><a href="/facts/Gryposaurus_latidens/mJwHvder">Gryposaurus latidens</a></i> and <i><a href="/facts/Hypacrosaurus/BeeBEEva">Hypacrosaurus</a></i> have been found to coexist with <i><a href="/facts/Maiasaura/6eqMzunc">Maiasaura</a></i>.<a class="footnote-ref" id="fnref:42" href="#fn:42"><sup>42</sup></a> Further, there are fossil teeth that seem to show the presence of certain taxa are unbroken throughout the whole formation.<a class="footnote-ref" id="fnref:43" href="#fn:43"><sup>43</sup></a> </p><p>Nevertheless, some true changes in faunal composition seem to occur in the upper Two Medicine. The appearance of <i>Maiasaura</i> in the formation precedes the arrival of a diverse variety of other ornithischians.<a class="footnote-ref" id="fnref:44" href="#fn:44"><sup>44</sup></a> According to David Trexler, thorough examination of strata found along the Two Medicine River (which exposes the entire upper half of the Two Medicine Formation) indicates that the apparent diversification was a real event rather than a result of preservational biases.<a class="footnote-ref" id="fnref:45" href="#fn:45"><sup>45</sup></a> </p><p>The timeline below follows the stratigraphic chart presented by Horner <i>et al.</i> 2001.<a class="footnote-ref" id="fnref:46" href="#fn:46"><sup>46</sup></a> </p> <h2 id="dinosaurs">Dinosaurs</h2> <p>Some of the dinosaurs from the formation have been speculated to show signs of drought related death.<a class="footnote-ref" id="fnref:47" href="#fn:47"><sup>47</sup></a> Very few articulated dinosaurs have been found in the formation; most specimens are isolated, bone bed, poorly preserved or broken remains.<a class="footnote-ref" id="fnref:48" href="#fn:48"><sup>48</sup></a> Early studies assumed that the Two Medicine Formation would have the same dinosaurs as the Judith River Formation.<a class="footnote-ref" id="fnref:49" href="#fn:49"><sup>49</sup></a> It was only in 1978, that it was discovered that the formation had endemic dinosaurs.<a class="footnote-ref" id="fnref:50" href="#fn:50"><sup>50</sup></a> Even some genera regarded as wide-ranging predators exhibited a species difference between the Two Medicine and other formations.<a class="footnote-ref" id="fnref:51" href="#fn:51"><sup>51</sup></a> No ecological barriers have been postulated apart from upland/lowland habitat preference differences between the Two Medicine and Judith River Formation.<a class="footnote-ref" id="fnref:52" href="#fn:52"><sup>52</sup></a> There is no unequivocal evidence for intermingling between the wildlife of the Two Medicine and geographically adjacent contemporary formations.<a class="footnote-ref" id="fnref:53" href="#fn:53"><sup>53</sup></a> Dinosaur remains are more common in the upper part of the Two Medicine.<a class="footnote-ref" id="fnref:54" href="#fn:54"><sup>54</sup></a> </p> <h3>Ankylosaurs</h3> <table><tbody><tr><th colspan="7"><a href="/facts/Ankylosaur/lXPkLf0e">Ankylosaurs</a> of the Two Medicine Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Edmontonia/1222bGhF">Edmontonia</a></i><a class="footnote-ref" id="fnref:55" href="#fn:55"><sup>55</sup></a></td><td><i>E. rugosidens</i><a class="footnote-ref" id="fnref:56" href="#fn:56"><sup>56</sup></a></td><td><ul><li>Landslide Butte</li><li>Two Medicine River</li></ul></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:57" href="#fn:57"><sup>57</sup></a></td><td>A skull with right mandible, cervical vertebrae, dorsal vertebrae, sacral vertebrae, caudal vertebrae, ribs, partial right ilium, left and right ischia, right pubis and osteoderms.</td><td>A <a href="/facts/Nodosaurid/SjMEQugE">nodosaurid</a> also known from the <a href="/facts/Horseshoe_Canyon_Formation/9qcE6fkz">Horseshoe Canyon Formation</a> and <a href="/facts/Dinosaur_Park_Formation/fJwQPd31">Dinosaur Park Formation</a>.</td><td></td></tr><tr><td rowspan="2"><i><a href="/facts/Euoplocephalus/px55n2Gk">Euoplocephalus</a></i><a class="footnote-ref" id="fnref:58" href="#fn:58"><sup>58</sup></a></td><td><i>E. tutus</i><a class="footnote-ref" id="fnref:59" href="#fn:59"><sup>59</sup></a></td><td></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:60" href="#fn:60"><sup>60</sup></a></td><td></td><td>Misclassified, actually represent <i><a href="/facts/Scolosaurus/J7SDTSMr">Scolosaurus</a></i></td><td></td></tr><tr><td>Indeterminate</td><td><ul><li>Landslide Butte</li><li>Two Medicine River</li></ul></td><td>Flag Butt Member</td><td></td><td>Misclassified, probably <i><a href="/facts/Scolosaurus/J7SDTSMr">Scolosaurus</a></i></td><td></td></tr><tr><td><i><a href="/facts/Oohkotokia/mzFmRyL7">Oohkotokia</a></i><a class="footnote-ref" id="fnref:61" href="#fn:61"><sup>61</sup></a></td><td><i>O. horneri</i><a class="footnote-ref" id="fnref:62" href="#fn:62"><sup>62</sup></a></td><td></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:63" href="#fn:63"><sup>63</sup></a></td><td>[Four] skulls, cervical vertebra, proximal caudal vertebra, ribs partial scapula, distal humerus, a left scapulocoracoid, ischia, [two] tail clubs, cervical half-ring, osteoderms, fragments and an undescribed specimen.</td><td>Penkalski (2013) referred to <i>Oohkotokia</i> all <a href="/facts/Ankylosaurine/6j7lP5gh">ankylosaurine</a> specimens from this formation.<a class="footnote-ref" id="fnref:64" href="#fn:64"><sup>64</sup></a> Arbour and Currie (2013) later referred <i>Oohkotokia</i> to <i>Scolosaurus</i>.<a class="footnote-ref" id="fnref:65" href="#fn:65"><sup>65</sup></a></td><td></td></tr><tr><td><i><a href="/facts/Scolosaurus/J7SDTSMr">Scolosaurus</a></i><a class="footnote-ref" id="fnref:66" href="#fn:66"><sup>66</sup></a></td><td><i>S. cutleri</i><a class="footnote-ref" id="fnref:67" href="#fn:67"><sup>67</sup></a></td><td></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:68" href="#fn:68"><sup>68</sup></a></td><td>[Four] skulls, cervical vertebra, proximal caudal vertebra, ribs partial scapula, distal humerus, a left scapulocoracoid, ischia, [two] tail clubs, cervical half-ring, osteoderms, fragments and an undescribed specimen.</td><td>An <a href="/facts/Ankylosaurine/6j7lP5gh">ankylosaurine</a> <a href="/facts/Ankylosaurid/RsePQquY">ankylosaurid</a>. These remains were previously considered to represent <i>Euoplocephalus</i> and then referred to <i>Oohkotokia</i> before being placed in <i>Scolosaurus</i>.</td><td></td></tr></tbody></table> <h3>Avialans</h3> <table><tbody><tr><th colspan="7"><a href="/facts/Aviala/RJF6zwLP">Avialans</a> of the Two Medicine Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Gettyia/N1qRYnqN">Gettyia</a></i><a class="footnote-ref" id="fnref:69" href="#fn:69"><sup>69</sup></a><a class="footnote-ref" id="fnref:70" href="#fn:70"><sup>70</sup></a></td><td><i>G. gloriae</i><a class="footnote-ref" id="fnref:71" href="#fn:71"><sup>71</sup></a></td><td></td><td>Upper<a class="footnote-ref" id="fnref:72" href="#fn:72"><sup>72</sup></a></td><td>"Tarsometatarsus"<a class="footnote-ref" id="fnref:73" href="#fn:73"><sup>73</sup></a></td><td>An <a href="/facts/Avisauridae/oyshRLro">avisaurid</a> <a href="/facts/Enantiornithean/kuC8WyFS">enantiornithean</a></td><td></td></tr></tbody></table> <h3>Ceratopsians</h3> <table><tbody><tr><th colspan="7"><a href="/facts/Ceratopsian/dQ1v7zFC">Ceratopsians</a> of the Two Medicine Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Achelousaurus/53hgrWNW">Achelousaurus</a></i><a class="footnote-ref" id="fnref:74" href="#fn:74"><sup>74</sup></a></td><td><i>A. horneri</i><a class="footnote-ref" id="fnref:75" href="#fn:75"><sup>75</sup></a></td><td>Landslide Butte</td><td>Upper Flag Butte Member<a class="footnote-ref" id="fnref:76" href="#fn:76"><sup>76</sup></a></td><td>"[Three] partial skulls, [one] partial skeleton."<a class="footnote-ref" id="fnref:77" href="#fn:77"><sup>77</sup></a></td><td>A <a href="/facts/Centrosaurine/TA93Tgk5">centrosaurine</a> <a href="/facts/Ceratopsid/H6zh5uAe">ceratopsid</a></td><td></td></tr><tr><td><i><a href="/facts/Brachyceratops/XouSbsrj">Brachyceratops</a></i><a class="footnote-ref" id="fnref:78" href="#fn:78"><sup>78</sup></a></td><td><i>B. montanensis</i><a class="footnote-ref" id="fnref:79" href="#fn:79"><sup>79</sup></a></td><td></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:80" href="#fn:80"><sup>80</sup></a></td><td>"[Six] partial skulls, skeletons, subadult."<a class="footnote-ref" id="fnref:81" href="#fn:81"><sup>81</sup></a></td><td>A centrosaurine ceratopsid. Might be a juvenile <i>Styracosaurus</i></td><td></td></tr><tr><td><i><a href="/facts/Cerasinops/xLSxHH7E">Cerasinops</a></i></td><td><i>C. hodgskissi</i><a class="footnote-ref" id="fnref:82" href="#fn:82"><sup>82</sup></a></td><td></td><td>Lower Hagans Crossing Member<a class="footnote-ref" id="fnref:83" href="#fn:83"><sup>83</sup></a></td><td></td><td>A <a href="/facts/Leptoceratopsid/keggzzpw">leptoceratopsid</a></td><td></td></tr><tr><td><i><a href="/facts/Einiosaurus/qWdVkW8T">Einiosaurus</a></i><a class="footnote-ref" id="fnref:84" href="#fn:84"><sup>84</sup></a></td><td><i>E. procurvicornis</i><a class="footnote-ref" id="fnref:85" href="#fn:85"><sup>85</sup></a></td><td>Landslide Butte</td><td>Upper Flag Butte Member<a class="footnote-ref" id="fnref:86" href="#fn:86"><sup>86</sup></a></td><td>"[Three] adult skulls, juvenile and subadult cranial and postcranial elements."<a class="footnote-ref" id="fnref:87" href="#fn:87"><sup>87</sup></a></td><td>A centrosaurine ceratopsid</td><td></td></tr><tr><td><i><a href="/facts/Prenoceratops/bjuuE8A7">Prenoceratops</a></i></td><td><i>P. pieganensis</i></td><td></td><td>Upper Flag Butte Member<a class="footnote-ref" id="fnref:88" href="#fn:88"><sup>88</sup></a></td><td></td><td>A leptoceratopsid</td><td></td></tr><tr><td><i><a href="/facts/Stellasaurus/3yN9qWRn">Stellasaurus</a></i><a class="footnote-ref" id="fnref:89" href="#fn:89"><sup>89</sup></a></td><td><i>S. ancellae</i></td><td>Landslide Butte</td><td>Flag Butte Member<a class="footnote-ref" id="fnref:90" href="#fn:90"><sup>90</sup></a></td><td>"Nasal horn and fragmentary parietal frill."</td><td>A centrosaurine ceratopsid</td><td></td></tr><tr><td><i><a href="/facts/Styracosaurus/UuQQoD6I">Styracosaurus</a></i><a class="footnote-ref" id="fnref:91" href="#fn:91"><sup>91</sup></a></td><td><i>S. ovatus</i></td><td></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:92" href="#fn:92"><sup>92</sup></a></td><td>Fragmentary parietal frill.</td><td>A centrosaurine ceratopsid</td><td></td></tr></tbody></table> <h3>Non-avialan eumaniraptorans</h3> <table><tbody><tr><th colspan="7">Non-<a href="/facts/Avialan/0296eLy8">avialan</a> <a href="/facts/Eumaniraptora/LB00ptV1">eumaniraptorans</a> of the Two Medicine Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Bambiraptor/VQxbq9Fq">Bambiraptor</a></i><a class="footnote-ref" id="fnref:93" href="#fn:93"><sup>93</sup></a></td><td><i>B. feinbergorum</i><a class="footnote-ref" id="fnref:94" href="#fn:94"><sup>94</sup></a></td><td></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:95" href="#fn:95"><sup>95</sup></a></td><td>"Almost complete skull and postcrania,"<a class="footnote-ref" id="fnref:96" href="#fn:96"><sup>96</sup></a> <a href="/facts/Type_specimen/mTM1beKX">type specimen</a></td><td>A <a href="/facts/Saurornitholestinae/qhuPFcrY">saurornitholestine</a> <a href="/facts/Dromaeosaur/mLeNBfBQ">dromaeosaur</a></td><td></td></tr><tr><td><i><a href="/facts/Dromaeosaurus/w9IaKwkw">Dromaeosaurus</a></i><a class="footnote-ref" id="fnref:97" href="#fn:97"><sup>97</sup></a></td><td>Indeterminate<a class="footnote-ref" id="fnref:98" href="#fn:98"><sup>98</sup></a></td><td></td><td>Hagans Crossing Member<a class="footnote-ref" id="fnref:99" href="#fn:99"><sup>99</sup></a></td><td></td><td></td><td></td></tr><tr><td><i><a href="/facts/Ricardoestesia/vWUz1BsJ">Richardoestesia</a></i><a class="footnote-ref" id="fnref:100" href="#fn:100"><sup>100</sup></a></td><td>Indeterminate<a class="footnote-ref" id="fnref:101" href="#fn:101"><sup>101</sup></a></td><td></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:102" href="#fn:102"><sup>102</sup></a></td><td>Teeth</td><td></td><td></td></tr><tr><td><i><a href="/facts/Saurornitholestes/TUXqUqWB">Saurornitholestes</a></i><a class="footnote-ref" id="fnref:103" href="#fn:103"><sup>103</sup></a></td><td>Indeterminate<a class="footnote-ref" id="fnref:104" href="#fn:104"><sup>104</sup></a></td><td><ul><li>"Choteau/Bynum"</li><li>Landslide Butte</li><li>Two Medicine River</li></ul></td><td><ul><li>Flag Butte Member<a class="footnote-ref" id="fnref:105" href="#fn:105"><sup>105</sup></a></li><li>Hagans Crossing Member<a class="footnote-ref" id="fnref:106" href="#fn:106"><sup>106</sup></a></li></ul></td><td>Partial skeleton, isolated pedal elements</td><td>A saurornitholestine dromaeosaur</td><td></td></tr><tr><td><i><a href="/facts/Troodon/cRJams94">Troodon</a></i><a class="footnote-ref" id="fnref:107" href="#fn:107"><sup>107</sup></a></td><td><i>formosus</i>?<a class="footnote-ref" id="fnref:108" href="#fn:108"><sup>108</sup></a></td><td></td><td>Flag Butte Member</td><td>Partial skulls, several vertebrae, ribs, gastralia, chevrons, a sacrum, partial pelvis, and partial fore and hind limbs</td><td>The remains are controversial, typically assigned to <i>Troodon</i>, but later studies suggested that the genus may be <a href="/facts/Nomen_dubium/pxPDpId7">dubious</a>. However, it has been proposed that these remains do indeed belong to <i>Troodon</i> formosus, and that one specimen should probably serve as the <a href="/facts/Neotype/mTM1beKX">neotype</a>.</td><td></td></tr></tbody></table> <h3>Ornithopods</h3> <p>An unidentified <a href="/facts/Lambeosaurine/spWvHLLV">lambeosaurine</a> has been collected from the same stratigraphic placement, west of Bynum, and is in preparation at The Montana Dinosaur Center <a class="footnote-ref" id="fnref:109" href="#fn:109"><sup>109</sup></a> </p> <table><tbody><tr><th colspan="7"><a href="/facts/Ornithopod/HpWlQsXI">Ornithopods</a> and <a href="/facts/Parksosauridae/jUqEevu1">parksosaurs</a> of the Two Medicine Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Acristavus/SvPmM8su">Acristavus</a></i><a class="footnote-ref" id="fnref:110" href="#fn:110"><sup>110</sup></a></td><td><i>A. gagslarsoni</i></td><td></td><td>Lower Hagans Crossing Member<a class="footnote-ref" id="fnref:111" href="#fn:111"><sup>111</sup></a></td><td></td><td>A <a href="/facts/Saurolophine/6bjxGDg9">saurolophine</a> <a href="/facts/Hadrosaur/v4t7JSPB">hadrosaur</a></td><td></td></tr><tr><td><i><a href="/facts/Glishades/NcGT9qKF">Glishades</a></i></td><td><i>G. ericksoni</i></td><td></td><td></td><td></td><td>A <a href="/facts/Hadrosauroid/FJ49JytU">hadrosauroid</a><a class="footnote-ref" id="fnref:112" href="#fn:112"><sup>112</sup></a> or an <a href="/facts/Nomen_dubium/pxPDpId7">indeterminate</a> juvenile saurolophine hadrosaur.<a class="footnote-ref" id="fnref:113" href="#fn:113"><sup>113</sup></a></td><td></td></tr><tr><td rowspan="2"><i><a href="/facts/Gryposaurus/mJwHvder">Gryposaurus</a></i><a class="footnote-ref" id="fnref:114" href="#fn:114"><sup>114</sup></a></td><td><i>G. latidens</i><a class="footnote-ref" id="fnref:115" href="#fn:115"><sup>115</sup></a></td><td>Two Medicine River</td><td><ul><li>Lower Hagans Crossing Member<a class="footnote-ref" id="fnref:116" href="#fn:116"><sup>116</sup></a></li></ul></td><td>"Several partial skulls and postcranial skeletons."<a class="footnote-ref" id="fnref:117" href="#fn:117"><sup>117</sup></a> Also known from isolated teeth which may have been redeposited fossils, although this explanation is unlikely.<a class="footnote-ref" id="fnref:118" href="#fn:118"><sup>118</sup></a></td><td>A saurolophine hadrosaur. The isolated <i>G. latidens</i> teeth are a rare component of channel lag deposits in the middle portion of the formation.<a class="footnote-ref" id="fnref:119" href="#fn:119"><sup>119</sup></a></td><td rowspan="2"></td></tr><tr><td>Indeterminate<a class="footnote-ref" id="fnref:120" href="#fn:120"><sup>120</sup></a></td><td></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:121" href="#fn:121"><sup>121</sup></a></td><td></td><td></td></tr><tr><td><i><a href="/facts/Hypacrosaurus/BeeBEEva">Hypacrosaurus</a></i><a class="footnote-ref" id="fnref:122" href="#fn:122"><sup>122</sup></a></td><td><i>H. stebingeri</i><a class="footnote-ref" id="fnref:123" href="#fn:123"><sup>123</sup></a></td><td><ul><li>Landslide Butte</li><li>Two Medicine River</li></ul></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:124" href="#fn:124"><sup>124</sup></a></td><td></td><td>A very abundant species of <a href="/facts/Lambeosaurine/spWvHLLV">lambeosaurine</a> hadrosaur.<a class="footnote-ref" id="fnref:125" href="#fn:125"><sup>125</sup></a></td><td></td></tr><tr><td><i><a href="/facts/Maiasaura/6eqMzunc">Maiasaura</a></i><a class="footnote-ref" id="fnref:126" href="#fn:126"><sup>126</sup></a></td><td><i>M. peeblesorum</i><a class="footnote-ref" id="fnref:127" href="#fn:127"><sup>127</sup></a></td><td><ul><li>"Choteau/Bynum"</li><li>Two Medicine River</li></ul></td><td>Upper Hagans Crossing Member<a class="footnote-ref" id="fnref:128" href="#fn:128"><sup>128</sup></a></td><td>"More than [two hundred] specimens including articulated skull and postcrania, embryo to adult."<a class="footnote-ref" id="fnref:129" href="#fn:129"><sup>129</sup></a></td><td>A saurolophine hadrosaur. <a href="/facts/Choteau/d1HYSBK9">Choteau</a> <i><a href="/facts/Maiasaura/6eqMzunc">Maiasaura</a></i> remains are found in higher strata than their Two Medicine River counterparts.<a class="footnote-ref" id="fnref:130" href="#fn:130"><sup>130</sup></a> It is the most common dinosaur found in the Egg Mountain locality.<a class="footnote-ref" id="fnref:131" href="#fn:131"><sup>131</sup></a></td><td></td></tr><tr><td><i><a href="/facts/Orodromeus/hJ7tFcFI">Orodromeus</a></i><a class="footnote-ref" id="fnref:132" href="#fn:132"><sup>132</sup></a></td><td><i>O. makelai</i><a class="footnote-ref" id="fnref:133" href="#fn:133"><sup>133</sup></a></td><td>"Choteau/Bynum"</td><td>Lacustrine Interval, Lower Flag Butte Member<a class="footnote-ref" id="fnref:134" href="#fn:134"><sup>134</sup></a></td><td></td><td>An <a href="/facts/Orodromine/F6F0CAo1">orodromine</a> <a href="/facts/Thescelosauridae/jUqEevu1">thescelosaur</a> which was the most common small herbivore in the Egg Mountain area.<a class="footnote-ref" id="fnref:135" href="#fn:135"><sup>135</sup></a><a class="footnote-ref" id="fnref:136" href="#fn:136"><sup>136</sup></a></td><td></td></tr><tr><td><i><a href="/facts/Prosaurolophus/qVyY99z3">Prosaurolophus</a></i><a class="footnote-ref" id="fnref:137" href="#fn:137"><sup>137</sup></a></td><td><i>P. maximus</i><a class="footnote-ref" id="fnref:138" href="#fn:138"><sup>138</sup></a></td><td><ul><li>Landslide Butte</li><li>Two Medicine River</li></ul></td><td>Upper Flag Butte Member<a class="footnote-ref" id="fnref:139" href="#fn:139"><sup>139</sup></a></td><td>"Disarticulated, associated skull and postcrania pertaining to at least [four] individuals."<a class="footnote-ref" id="fnref:140" href="#fn:140"><sup>140</sup></a></td><td>A saurolophine hadrosaur. <i>Prosaurolophus blackfeetensis</i>, erected for Two Medicine fossils, is a synonym of <i>P. maximus</i>.<a class="footnote-ref" id="fnref:141" href="#fn:141"><sup>141</sup></a></td><td></td></tr></tbody></table> <h3>Oviraptorosaurs</h3> <p>The first find of an oviraptorosaur in Montana was an <a href="/facts/Articular/mtMzom1h">articular</a> region from the lower jaw of <i><a href="/facts/Caenagnathus_sternbergi/Q5tsabfD">Caenagnathus sternbergi</a></i>, from the Two Medicine Formation, according to a 2001 paper by David J. Varrichio.<a class="footnote-ref" id="fnref:142" href="#fn:142"><sup>142</sup></a> This species had previously only been known from the Canadian province of <a href="/facts/Alberta/PjKJFbJY">Alberta</a>.<a class="footnote-ref" id="fnref:143" href="#fn:143"><sup>143</sup></a> Varricchio observes that during the late <a href="/facts/Campanian/MoPSxcqL">Campanian</a>, Alberta and Montana had very similar <a href="/facts/Theropods/KUzy1Ef7">theropods</a> despite significant differences in the types of herbivorous dinosaur faunas.<a class="footnote-ref" id="fnref:144" href="#fn:144"><sup>144</sup></a> </p> <table><tbody><tr><th colspan="7"><a href="/facts/Oviraptorosaurs/C6puS7KW">Oviraptorosaurs</a> of the Two Medicine Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Chirostenotes/Q5tsabfD">Chirostenotes</a></i><a class="footnote-ref" id="fnref:145" href="#fn:145"><sup>145</sup></a></td><td><i>C. pergracilis</i><a class="footnote-ref" id="fnref:146" href="#fn:146"><sup>146</sup></a></td><td></td><td></td><td>Known from the articular region of a lower jaw, catalogued as MOR 1107<a class="footnote-ref" id="fnref:147" href="#fn:147"><sup>147</sup></a></td><td>Was previously referred to <i>Caenagnathus sternbergi</i> (a synonym of <i>Chirostenotes)</i></td><td></td></tr></tbody></table> <h3>Tyrannosauroids</h3> <table><tbody><tr><th colspan="7"><a href="/facts/Tyrannosauroid/sIUowTnq">Tyrannosauroids</a> of the Two Medicine Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td rowspan="2"><i><a href="/facts/Daspletosaurus/fccLgNmd">Daspletosaurus</a></i><a class="footnote-ref" id="fnref:148" href="#fn:148"><sup>148</sup></a></td><td><i>D. horneri</i><a class="footnote-ref" id="fnref:149" href="#fn:149"><sup>149</sup></a></td><td></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:150" href="#fn:150"><sup>150</sup></a></td><td>Bonebed<a class="footnote-ref" id="fnref:151" href="#fn:151"><sup>151</sup></a></td><td></td><td></td></tr><tr><td><i>D. wilsoni</i></td><td></td><td>Flag Butte Member</td><td>Bonebed<a class="footnote-ref" id="fnref:152" href="#fn:152"><sup>152</sup></a></td><td></td><td></td></tr><tr><td><i><a href="/facts/Gorgosaurus/jTP8bUS3">Gorgosaurus</a></i></td><td><i>G. sp</i></td><td>"Choteau/Bynum"</td><td>Flag Butte Member</td><td></td><td>Includes TCM 2001.89.1., the undescribed skeleton</td><td></td></tr><tr><td><a href="/facts/Teratophoneini/EBAte1P1">Teratophoneini</a></td><td>Indeterminate</td><td></td><td></td><td>Frontal<a class="footnote-ref" id="fnref:153" href="#fn:153"><sup>153</sup></a></td><td>Phylogenetic analysis recovers it in Teratophoneini<a class="footnote-ref" id="fnref:154" href="#fn:154"><sup>154</sup></a></td><td></td></tr><tr><td><i><a href="/facts/Tyrannosaurus/HQw1cywX">Tyrannosaurus</a></i></td><td><i>T.</i> sp.</td><td></td><td>Flag Butte Member<a class="footnote-ref" id="fnref:155" href="#fn:155"><sup>155</sup></a></td><td>right lacrimal, isolated tooth<a class="footnote-ref" id="fnref:156" href="#fn:156"><sup>156</sup></a><a class="footnote-ref" id="fnref:157" href="#fn:157"><sup>157</sup></a></td><td>More similar to Tyrannosaurini than other Campanian tyrannosaurids<a class="footnote-ref" id="fnref:158" href="#fn:158"><sup>158</sup></a></td><td></td></tr></tbody></table> <table><tbody><tr><th></th><th></th><th></th></tr><tr><td>Color key<table><tbody><tr><td><a href="/facts/Taxon/caQC1e87">Taxon</a></td><td>Reclassified taxon</td><td>Taxon falsely reported as present</td><td>Dubious taxon or junior synonym</td><td><a href="/facts/Ichnotaxon/edWeWT20">Ichnotaxon</a></td><td><a href="/facts/Ootaxon/nH5XHNRS">Ootaxon</a></td><td><a href="/facts/Morphotaxon/zWsaRMUe">Morphotaxon</a></td></tr></tbody></table></td><td></td><td>Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.</td></tr></tbody></table> <h2 id="other-fauna">Other fauna</h2> <p>Many other fossil animals have been found, such as freshwater <a href="/facts/Bivalves/IQDxHVc8">bivalves</a>, <a href="/facts/Gastropod/rYPqm3PT">gastropods</a>, <a href="/facts/Turtle/X0ImnxYt">turtles</a>, lizards such as <i><a href="/facts/Magnuviator/U122084t">Magnuviator</a></i>, and <a href="/facts/Choristodera/rTolEzdL">champsosaurs</a>. The multituberculate mammal <i><a href="/facts/Cimexomys/tjzVF1L6">Cimexomys</a></i> has been found on Egg Mountain. The species <i><a href="/facts/Piksi/Ht8ek8qM">Piksi barbarulna</a></i> was described based on forelimb bones from the Two Medicine Formation; it was initially thought to be a <a href="/facts/Bird/RJF6zwLP">bird</a>, but subsequently it was reinterpreted as a <a href="/facts/Pterosaur/YNQwa1ru">pterosaur</a>, likely a member of <a href="/facts/Ornithocheiroidea/hfQ9YPyI">Ornithocheiroidea</a>.<a class="footnote-ref" id="fnref:159" href="#fn:159"><sup>159</sup></a> <a href="/facts/Azhdarchoidea/W72cQxog">Azhdarchoid</a> pterosaurs are also known from the Two Medicine Formation, including a very large, yet-unnamed <a href="/facts/Azhdarchidae/km8IDnS4">azhdarchid</a>, the estimated wingspan of which was 8 metres (26 ft), and smaller <i><a href="/facts/Montanazhdarcho_minor/afGbALjP">Montanazhdarcho minor</a></i>, a non-azhdarchid azhdarchoid.<a class="footnote-ref" id="fnref:160" href="#fn:160"><sup>160</sup></a><a class="footnote-ref" id="fnref:161" href="#fn:161"><sup>161</sup></a> <a href="/facts/Insect/IgDNvnUL">Insect</a> and <a href="/facts/Mammal/phSwTzBo">mammal</a> <a href="/facts/Burrow/vgY0GpEH">burrows</a> have also been discovered, as well as dinosaur <a href="/facts/Coprolites/VX4K6BNX">coprolites</a>. </p> <h2 id="see-also">See also</h2> <ul><li><a href="/facts/List_of_dinosaur-bearing_rock_formations/G8KXXsw0">List of dinosaur-bearing rock formations</a></li> <li><a href="/facts/List_of_fossil_sites/6gYhEPes">List of fossil sites</a> <i>(with link directory)</i></li></ul> <h3>Bibliography</h3> <ul><li>Dodson, P., C.A. Forster, and S.D. Sampson. 2004. Ceratopsidae in Weishampel, D.B., P. Dodson, and H. Osmolska (eds.) <i>The Dinosauria</i>. 2nd Edition, University of California Press.</li> <li>Rogers, R.R. (1990). "Taphonomy of three dinosaur bone beds in the Upper Cretaceous Two Medicine Formation of northwestern Montana: evidence for drought-related mortality". <i>PALAIOS</i>. 5 (5): 394–413. <a href="/facts/Bibcode_(identifier)/9HtdQSGB">Bibcode</a>:<a href="https://ui.adsabs.harvard.edu/abs/1990Palai...5..394R">1990Palai...5..394R</a>. <a href="/facts/Doi_(identifier)/muM9Etpq">doi</a>:<a href="https://doi.org/10.2307%2F3514834">10.2307/3514834</a>. <a href="/facts/JSTOR_(identifier)/YTeVmaJ7">JSTOR</a> <a href="https://www.jstor.org/stable/3514834">3514834</a>.</li> <li>Trexler, D., 2001, Two Medicine Formation, Montana: geology and fauna: In: <i>Mesozoic Vertebrate Life</i>, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, pp. 298–309.</li> <li>Varricchio, D.J. (1995). <a href="http://doc.rero.ch/record/14441/files/PAL_E1625.pdf">"Taphonomy of Jack's Birthday Site, a diverse dinosaur bonebed from the Upper Cretaceous Two Medicine Formation of Montana"</a> (PDF). <i>Palaeogeography, Palaeoclimatology, Palaeoecology</i>. 114 (2–4): 297–323. <a href="/facts/Bibcode_(identifier)/9HtdQSGB">Bibcode</a>:<a href="https://ui.adsabs.harvard.edu/abs/1995PPP...114..297V">1995PPP...114..297V</a>. <a href="/facts/Doi_(identifier)/muM9Etpq">doi</a>:<a href="https://doi.org/10.1016%2F0031-0182%2894%2900084-l">10.1016/0031-0182(94)00084-l</a>.</li> <li>Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropoda) dinosaurs from Montana. pp. 42–57 in D. H. Tanke and K. Carpenter (eds.), Mesozoic Vertebrate Life. Indiana University Press, Indianapolis.</li> <li><a href="/facts/David_B._Weishampel/j6GTWKnC">Weishampel</a>, David B.; <a href="/facts/Peter_Dodson/zuCv5evX">Dodson</a>, Peter; <a href="/facts/Halszka_Osm%25C3%25B3lska/kF2xGfyv">Osmólska</a>, Halszka, eds. (2004). <a href="https://books.google.com/books?id=vtZFDb_iw40C"><i>The Dinosauria, 2nd edition</i></a>. Berkeley: University of California Press. pp. 1–880. <a href="/facts/ISBN_(identifier)/15AdSPa9">ISBN</a> 0-520-24209-2. Retrieved 2019-02-21.</li></ul> <h2 id="references">References</h2> <ol> <li id="fn:1"><p>"Previous Work". Trexler (2001); page 300. <a href="#fnref:1" class="footnote-back-ref">↩</a></p></li> <li id="fn:2"><p>"Previous Work". Trexler (2001); page 300. <a href="#fnref:2" class="footnote-back-ref">↩</a></p></li> <li id="fn:3"><p>"Previous Work". Trexler (2001); page 300. <a href="#fnref:3" class="footnote-back-ref">↩</a></p></li> <li id="fn:4"><p>"Previous Work". Trexler (2001); page 300. <a href="#fnref:4" class="footnote-back-ref">↩</a></p></li> <li id="fn:5"><p>"Previous Work". Trexler (2001); page 300. <a href="#fnref:5" class="footnote-back-ref">↩</a></p></li> <li id="fn:6"><p>"Previous Work". Trexler (2001); page 300. <a href="#fnref:6" class="footnote-back-ref">↩</a></p></li> <li id="fn:7"><p>"Previous Work". Trexler (2001); page 300. <a href="#fnref:7" class="footnote-back-ref">↩</a></p></li> <li id="fn:8"><p>"Previous Work". Trexler (2001); page 300. <a href="#fnref:8" class="footnote-back-ref">↩</a></p></li> <li id="fn:9"><p>"Previous Work". Trexler (2001); page 300. <a href="#fnref:9" class="footnote-back-ref">↩</a></p></li> <li id="fn:10"><p>"Introduction". Trexler (2001); pages 299-300. <a href="#fnref:10" class="footnote-back-ref">↩</a></p></li> <li id="fn:11"><p>"Introduction". Trexler (2001); page 299. <a href="#fnref:11" class="footnote-back-ref">↩</a></p></li> <li id="fn:12"><p>"Introduction". Trexler (2001); pages 298-299. <a href="#fnref:12" class="footnote-back-ref">↩</a></p></li> <li id="fn:13"><p>Rogers, Raymond R.; Horner, John R.; Ramezani, Jahandar; Roberts, Eric M.; Varricchio, David J. (2024). "Updating the Upper Cretaceous (Campanian) Two Medicine Formation of Montana: Lithostratigraphic revisions, new CA-ID-TIMS U-Pb ages, and a calibrated framework for dinosaur occurrences". Geological Society of America Bulletin. doi:10.1130/B37498.1. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:13" class="footnote-back-ref">↩</a></p></li> <li id="fn:14"><p>"Geological Setting". Trexler (2001); pages 300-301. <a href="#fnref:14" class="footnote-back-ref">↩</a></p></li> <li id="fn:15"><p>"Geological Setting". Trexler (2001); page 301. <a href="#fnref:15" class="footnote-back-ref">↩</a></p></li> <li id="fn:16"><p>"Geological Setting". Trexler (2001); page 301. <a href="#fnref:16" class="footnote-back-ref">↩</a></p></li> <li id="fn:17"><p>"Geological Setting". Trexler (2001); page 301. <a href="#fnref:17" class="footnote-back-ref">↩</a></p></li> <li id="fn:18"><p>"Geological Setting". Trexler (2001); page 301. <a href="#fnref:18" class="footnote-back-ref">↩</a></p></li> <li id="fn:19"><p>"Geological Setting". Trexler (2001); page 301. <a href="#fnref:19" class="footnote-back-ref">↩</a></p></li> <li id="fn:20"><p>"Geological Setting". Trexler (2001); page 302. <a href="#fnref:20" class="footnote-back-ref">↩</a></p></li> <li id="fn:21"><p>"Geological Setting". Trexler (2001); page 302. <a href="#fnref:21" class="footnote-back-ref">↩</a></p></li> <li id="fn:22"><p>"Geological Setting". Trexler (2001); page 302. <a href="#fnref:22" class="footnote-back-ref">↩</a></p></li> <li id="fn:23"><p>"Geological Setting". Trexler (2001); page 302. <a href="#fnref:23" class="footnote-back-ref">↩</a></p></li> <li id="fn:24"><p>"Geological Setting". Trexler (2001); page 302. <a href="#fnref:24" class="footnote-back-ref">↩</a></p></li> <li id="fn:25"><p>"Geological Setting". Trexler (2001); page 302. <a href="#fnref:25" class="footnote-back-ref">↩</a></p></li> <li id="fn:26"><p>"Introduction". Trexler (2001); page 299. <a href="#fnref:26" class="footnote-back-ref">↩</a></p></li> <li id="fn:27"><p>"Introduction". Trexler (2001); page 299. <a href="#fnref:27" class="footnote-back-ref">↩</a></p></li> <li id="fn:28"><p>"Introduction". Trexler (2001); page 299. <a href="#fnref:28" class="footnote-back-ref">↩</a></p></li> <li id="fn:29"><p>"Geological Setting". Trexler (2001); page 302. <a href="#fnref:29" class="footnote-back-ref">↩</a></p></li> <li id="fn:30"><p>"Geological Setting". Trexler (2001); page 302. <a href="#fnref:30" class="footnote-back-ref">↩</a></p></li> <li id="fn:31"><p>"Geological Setting". Trexler (2001); page 301. <a href="#fnref:31" class="footnote-back-ref">↩</a></p></li> <li id="fn:32"><p>"Geological Setting". Trexler (2001); page 301. <a href="#fnref:32" class="footnote-back-ref">↩</a></p></li> <li id="fn:33"><p>"Geological Setting". Trexler (2001); page 302. <a href="#fnref:33" class="footnote-back-ref">↩</a></p></li> <li id="fn:34"><p>"Geological Setting". Trexler (2001); page 302. <a href="#fnref:34" class="footnote-back-ref">↩</a></p></li> <li id="fn:35"><p>"Abstract". Trexler (2001); page 298. <a href="#fnref:35" class="footnote-back-ref">↩</a></p></li> <li id="fn:36"><p>"Abstract". Trexler (2001); page 298. <a href="#fnref:36" class="footnote-back-ref">↩</a></p></li> <li id="fn:37"><p>"Abstract". Trexler (2001); page 298. <a href="#fnref:37" class="footnote-back-ref">↩</a></p></li> <li id="fn:38"><p>"Faunal Turnover, Migration, and Evolution". Trexler (2001); page 306. <a href="#fnref:38" class="footnote-back-ref">↩</a></p></li> <li id="fn:39"><p>"Faunal Turnover, Migration, and Evolution". Trexler (2001); page 304. <a href="#fnref:39" class="footnote-back-ref">↩</a></p></li> <li id="fn:40"><p>"Faunal Turnover, Migration, and Evolution". Trexler (2001); page 306. <a href="#fnref:40" class="footnote-back-ref">↩</a></p></li> <li id="fn:41"><p>"Two Medicine Fauna". Trexler (2001); page 303. <a href="#fnref:41" class="footnote-back-ref">↩</a></p></li> <li id="fn:42"><p>"Faunal Turnover, Migration, and Evolution". Trexler (2001); page 306. <a href="#fnref:42" class="footnote-back-ref">↩</a></p></li> <li id="fn:43"><p>"Faunal Turnover, Migration, and Evolution". Trexler (2001); page 306. <a href="#fnref:43" class="footnote-back-ref">↩</a></p></li> <li id="fn:44"><p>"Faunal Turnover, Migration, and Evolution". Trexler (2001); page 306. <a href="#fnref:44" class="footnote-back-ref">↩</a></p></li> <li id="fn:45"><p>"Faunal Turnover, Migration, and Evolution". Trexler (2001); page 306. <a href="#fnref:45" class="footnote-back-ref">↩</a></p></li> <li id="fn:46"><p>Horner, J. R., Schmitt, J. G., Jackson, F., & Hanna, R. (2001). Bones and rocks of the Upper Cretaceous Two Medicine-Judith River clastic wedge complex, Montana. In Field trip guidebook, Society of Vertebrate Paleontology 61st Annual Meeting: Mesozoic and Cenozoic Paleontology in the Western Plains and Rocky Mountains. Museum of the Rockies Occasional Paper (Vol. 3, pp. 3-14). <a href="#fnref:46" class="footnote-back-ref">↩</a></p></li> <li id="fn:47"><p>"Geological Setting". Trexler (2001); page 302. <a href="#fnref:47" class="footnote-back-ref">↩</a></p></li> <li id="fn:48"><p>"Two Medicine Fauna". Trexler (2001); page 302. <a href="#fnref:48" class="footnote-back-ref">↩</a></p></li> <li id="fn:49"><p>"Two Medicine Fauna". Trexler (2001); page 302. <a href="#fnref:49" class="footnote-back-ref">↩</a></p></li> <li id="fn:50"><p>"Two Medicine Fauna". Trexler (2001); page 302. <a href="#fnref:50" class="footnote-back-ref">↩</a></p></li> <li id="fn:51"><p>"Two Medicine Fauna". Trexler (2001); pages 302-303. <a href="#fnref:51" class="footnote-back-ref">↩</a></p></li> <li id="fn:52"><p>"Two Medicine Fauna". Trexler (2001); page 303. <a href="#fnref:52" class="footnote-back-ref">↩</a></p></li> <li id="fn:53"><p>"Two Medicine Fauna". Trexler (2001); page 303. <a href="#fnref:53" class="footnote-back-ref">↩</a></p></li> <li id="fn:54"><p>"Two Medicine Fauna". Trexler (2001); page 303. <a href="#fnref:54" class="footnote-back-ref">↩</a></p></li> <li id="fn:55"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:55" class="footnote-back-ref">↩</a></p></li> <li id="fn:56"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:56" class="footnote-back-ref">↩</a></p></li> <li id="fn:57"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:57" class="footnote-back-ref">↩</a></p></li> <li id="fn:58"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:58" class="footnote-back-ref">↩</a></p></li> <li id="fn:59"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:59" class="footnote-back-ref">↩</a></p></li> <li id="fn:60"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:60" class="footnote-back-ref">↩</a></p></li> <li id="fn:61"><p>Penkalski, P. (2013). "A new ankylosaurid from the late Cretaceous Two Medicine Formation of Montana, USA". Acta Palaeontologica Polonica. 59 (3): 617–634. doi:10.4202/app.2012.0125. <a href="https://doi.org/10.4202%2Fapp.2012.0125" target="_blank">https://doi.org/10.4202%2Fapp.2012.0125</a> <a href="#fnref:61" class="footnote-back-ref">↩</a></p></li> <li id="fn:62"><p>Penkalski, P. (2013). "A new ankylosaurid from the late Cretaceous Two Medicine Formation of Montana, USA". Acta Palaeontologica Polonica. 59 (3): 617–634. doi:10.4202/app.2012.0125. <a href="https://doi.org/10.4202%2Fapp.2012.0125" target="_blank">https://doi.org/10.4202%2Fapp.2012.0125</a> <a href="#fnref:62" class="footnote-back-ref">↩</a></p></li> <li id="fn:63"><p>Rogers, Raymond R.; Horner, John R.; Ramezani, Jahandar; Roberts, Eric M.; Varricchio, David J. (2024). "Updating the Upper Cretaceous (Campanian) Two Medicine Formation of Montana: Lithostratigraphic revisions, new CA-ID-TIMS U-Pb ages, and a calibrated framework for dinosaur occurrences". Geological Society of America Bulletin. doi:10.1130/B37498.1. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:63" class="footnote-back-ref">↩</a></p></li> <li id="fn:64"><p>Penkalski, P. (2013). "A new ankylosaurid from the late Cretaceous Two Medicine Formation of Montana, USA". Acta Palaeontologica Polonica. 59 (3): 617–634. doi:10.4202/app.2012.0125. <a href="https://doi.org/10.4202%2Fapp.2012.0125" target="_blank">https://doi.org/10.4202%2Fapp.2012.0125</a> <a href="#fnref:64" class="footnote-back-ref">↩</a></p></li> <li id="fn:65"><p>Arbour, Victoria M.; Currie, Philip J. (2013-05-08). "Euoplocephalus tutus and the Diversity of Ankylosaurid Dinosaurs in the Late Cretaceous of Alberta, Canada, and Montana, USA". PLOS ONE. 8 (5): e62421. Bibcode:2013PLoSO...862421A. doi:10.1371/journal.pone.0062421. ISSN 1932-6203. PMC 3648582. PMID 23690940. <a href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3648582" target="_blank">https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3648582</a> <a href="#fnref:65" class="footnote-back-ref">↩</a></p></li> <li id="fn:66"><p>Arbour, Victoria M.; Currie, Philip J. (8 May 2013). "Euoplocephalus tutus and the Diversity of Ankylosaurid Dinosaurs in the Late Cretaceous of Alberta, Canada, and Montana, USA". PLOS ONE. 8 (5): e62421. Bibcode:2013PLoSO...862421A. doi:10.1371/journal.pone.0062421. PMC 3648582. PMID 23690940. <a href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3648582" target="_blank">https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3648582</a> <a href="#fnref:66" class="footnote-back-ref">↩</a></p></li> <li id="fn:67"><p>Arbour, Victoria M.; Currie, Philip J. (8 May 2013). "Euoplocephalus tutus and the Diversity of Ankylosaurid Dinosaurs in the Late Cretaceous of Alberta, Canada, and Montana, USA". PLOS ONE. 8 (5): e62421. Bibcode:2013PLoSO...862421A. doi:10.1371/journal.pone.0062421. PMC 3648582. PMID 23690940. <a href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3648582" target="_blank">https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3648582</a> <a href="#fnref:67" class="footnote-back-ref">↩</a></p></li> <li id="fn:68"><p>Rogers, Raymond R.; Horner, John R.; Ramezani, Jahandar; Roberts, Eric M.; Varricchio, David J. (2024). "Updating the Upper Cretaceous (Campanian) Two Medicine Formation of Montana: Lithostratigraphic revisions, new CA-ID-TIMS U-Pb ages, and a calibrated framework for dinosaur occurrences". Geological Society of America Bulletin. doi:10.1130/B37498.1. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:68" class="footnote-back-ref">↩</a></p></li> <li id="fn:69"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:69" class="footnote-back-ref">↩</a></p></li> <li id="fn:70"><p>Atterholt, Jessie; Hutchison, J. Howard; O’Connor, Jingmai K. (2018). "The most complete enantiornithine from North America and a phylogenetic analysis of the Avisauridae". PeerJ. 6: e5910. doi:10.7717/peerj.5910. PMC 6238772. PMID 30479894. <a href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6238772" target="_blank">https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6238772</a> <a href="#fnref:70" class="footnote-back-ref">↩</a></p></li> <li id="fn:71"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:71" class="footnote-back-ref">↩</a></p></li> <li id="fn:72"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:72" class="footnote-back-ref">↩</a></p></li> <li id="fn:73"><p>"Table 11.1". in Weishampel, et al. (2004). Page 212. <a href="#fnref:73" class="footnote-back-ref">↩</a></p></li> <li id="fn:74"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:74" class="footnote-back-ref">↩</a></p></li> <li id="fn:75"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:75" class="footnote-back-ref">↩</a></p></li> <li id="fn:76"><p>Rogers, Raymond R.; Horner, John R.; Ramezani, Jahandar; Roberts, Eric M.; Varricchio, David J. (2024). "Updating the Upper Cretaceous (Campanian) Two Medicine Formation of Montana: Lithostratigraphic revisions, new CA-ID-TIMS U-Pb ages, and a calibrated framework for dinosaur occurrences". Geological Society of America Bulletin. doi:10.1130/B37498.1. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:76" class="footnote-back-ref">↩</a></p></li> <li id="fn:77"><p>"Table 23.1". in Weishampel, et al. (2004). Page 495. <a href="#fnref:77" class="footnote-back-ref">↩</a></p></li> <li id="fn:78"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:78" class="footnote-back-ref">↩</a></p></li> <li id="fn:79"><p>"3.11 Montana, United States; 6. Upper Two Medicine Formation". in Weishampel, et al. (2004). Page 583. <a href="#fnref:79" class="footnote-back-ref">↩</a></p></li> <li id="fn:80"><p>Rogers, Raymond R.; Horner, John R.; Ramezani, Jahandar; Roberts, Eric M.; Varricchio, David J. (2024). "Updating the Upper Cretaceous (Campanian) Two Medicine Formation of Montana: Lithostratigraphic revisions, new CA-ID-TIMS U-Pb ages, and a calibrated framework for dinosaur occurrences". 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