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Cobitis
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<h2 id="species">Species</h2> <p>There are currently 97 recognized species in this genus: </p> <ul><li><i>Cobitis amphilekta</i> Vasil'eva & Vasil'ev, 2012 (Khvalyn spined loach)<a class="footnote-ref" id="fnref:3" href="#fn:3"><sup>3</sup></a></li> <li><i><a href="/facts/Cobitis_arachthosensis/eBJ830ku">Cobitis arachthosensis</a></i> Economidis & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1996</li> <li><i>Cobitis arenae</i> (S. Y. Lin, 1934)</li> <li><i>Cobitis australis</i> Y. X. Chen, Y. F. Chen & D. K. He, 2013<a class="footnote-ref" id="fnref:4" href="#fn:4"><sup>4</sup></a></li> <li><i>Cobitis avicennae</i> Mousavi-Sabet, Vatandoust, Esmaeili, Geiger & <a href="/facts/J%25C3%25B6rg_Arthur_Freyhof/7MtHCSIz">Freyhof</a>, 2015 (Zagros spined loach)<a class="footnote-ref" id="fnref:5" href="#fn:5"><sup>5</sup></a></li> <li><i>Cobitis baishagensis</i> Y. X. Chen, X. Y. Sui, N. Liang & Y. F. Chen, 2015<a class="footnote-ref" id="fnref:6" href="#fn:6"><sup>6</sup></a></li> <li><i>Cobitis battalgilae</i> <a href="/facts/Mihai_C._B%25C4%2583cescu/0dMADKyp">Băcescu</a>, 1962 (Battalgil's spined loach)</li> <li><i><a href="/facts/Cobitis_bilineata/E12b4T9g">Cobitis bilineata</a></i> <a href="/facts/Giovanni_Canestrini/ba4Mrsfg">Canestrini</a>, 1865</li> <li><i><a href="/facts/Cobitis_bilseli/rLfwVGNY">Cobitis bilseli</a></i> <a href="/facts/Fahire_Battalgazi/Tt0oRIGd">Battalgil</a>, 1942 (Beyşehir spined loach)</li> <li><i><a href="/facts/Cobitis_biwae/JbgfDlzm">Cobitis biwae</a></i> <a href="/facts/David_Starr_Jordan/qLNo5UIR">D. S. Jordan</a> & <a href="/facts/John_Otterbein_Snyder/6qJ2qBPM">Snyder</a>, 1901</li> <li><i><a href="/facts/Cobitis_brevifasciata/jePQUqrU">Cobitis brevifasciata</a></i> (I. S. Kim & W. O. Lee, 1995)<a class="footnote-ref" id="fnref:7" href="#fn:7"><sup>7</sup></a></li> <li><i><a href="/facts/Cobitis_calderoni/W5JoA6nl">Cobitis calderoni</a></i> <a href="/facts/Mihai_C._B%25C4%2583cescu/0dMADKyp">Băcescu</a>, 1962</li> <li><i>Cobitis choii</i> I. S. Kim & Y. M. Son, 1984 (Choi's spined loach)</li> <li><i>Cobitis crassicauda</i> Y. X. Chen & Y. F. Chen, 2013<a class="footnote-ref" id="fnref:8" href="#fn:8"><sup>8</sup></a></li> <li><i><a href="/facts/Cobitis_dalmatina/cDXkTTe9">Cobitis dalmatina</a></i> <a href="/facts/Stanko_Luka_Karaman/vIDSfl02">S. L. Karaman</a>, 1928</li> <li><i>Cobitis damlae</i> Erk' Akan & Özdemir, 2014 (Dalaman spined loach)<a class="footnote-ref" id="fnref:9" href="#fn:9"><sup>9</sup></a></li> <li><i>Cobitis delicata</i> Niwa, 1937<a class="footnote-ref" id="fnref:10" href="#fn:10"><sup>10</sup></a></li> <li><i>Cobitis dolichorhynchus</i> <a href="/facts/John_Treadwell_Nichols/m3KgkrpI">Nichols</a>, 1918</li> <li><i>Cobitis dorademiri</i> Erk' Akan, Özdemir & Özeren, 2017<a class="footnote-ref" id="fnref:11" href="#fn:11"><sup>11</sup></a></li> <li><i>Cobitis elazigensis</i> Coad & Sarıeyyüpoglu, 1988 (Tigris spined loach)</li> <li><i><a href="/facts/Cobitis_elongata/Av2RGeWv">Cobitis elongata</a></i> <a href="/facts/Johann_Jakob_Heckel/tjkqX0qp">Heckel</a> & <a href="/facts/Rudolf_Kner/C4Y9kTWH">Kner</a>, 1857</li> <li><i>Cobitis elongatoides</i> <a href="/facts/Mihai_C._B%25C4%2583cescu/0dMADKyp">Băcescu</a> & R. F. Mayer, 1969</li> <li><i>Cobitis evreni</i> Erk' Akan, Özeren & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 2008 (Ceyhan spined loach)</li> <li><i><a href="/facts/Cobitis_fahireae/Qva139Ec">Cobitis fahireae</a></i> Erk' Akan, Ekmekçi & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1998 (Küçük Menderes spined loach)</li> <li><i>Cobitis faridpaki</i> Mousavi-Sabet, Vasil'eva, Vatandoust & Vasil'ev, 2011<a class="footnote-ref" id="fnref:12" href="#fn:12"><sup>12</sup></a></li> <li><i>Cobitis fasciola</i> Y. X. Chen & Y. F. Chen, 2013<a class="footnote-ref" id="fnref:13" href="#fn:13"><sup>13</sup></a></li> <li><i>Cobitis gladkovi</i> Vasil'ev & Vasil'eva, 2008</li> <li><i>Cobitis gracilis</i> Y. X. Chen & Y. F. Chen, 2016<a class="footnote-ref" id="fnref:14" href="#fn:14"><sup>14</sup></a></li> <li><i>Cobitis guttatus</i> (V. H. Nguyễn, 2005)</li> <li><i>Cobitis hankugensis</i> I. S. Kim, J. Y. Park, Y. M. Son & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 2003</li> <li><i><a href="/facts/Cobitis_hellenica/1S1g26Lu">Cobitis hellenica</a></i> Economidis & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1996</li> <li><i>Cobitis hereromacula</i> Y. X. Chen, X. Y. Sui, N. Liang & Y. F. Chen, 2015<a class="footnote-ref" id="fnref:15" href="#fn:15"><sup>15</sup></a></li> <li><i>Cobitis herzegoviniensis</i> Buj & Šanda, 2014<a class="footnote-ref" id="fnref:16" href="#fn:16"><sup>16</sup></a></li> <li><i>Cobitis hugowolfeldi</i> (<a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1993)<a class="footnote-ref" id="fnref:17" href="#fn:17"><sup>17</sup></a></li> <li><i>Cobitis illyrica</i> <a href="/facts/J%25C3%25B6rg_Arthur_Freyhof/7MtHCSIz">Freyhof</a> & Stelbrink, 2007</li> <li><i>Cobitis jadovaensis</i> Mustafić & Mrakovčić, 2008</li> <li><i>Cobitis kaibarai</i> Nakajima, 2012<a class="footnote-ref" id="fnref:18" href="#fn:18"><sup>18</sup></a></li> <li><i>Cobitis kellei</i> Erk' Akan, Ekmekçi & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1998 (Göksu spined loach)</li> <li><i>Cobitis koreensis</i> I. S. Kim, 1975<a class="footnote-ref" id="fnref:19" href="#fn:19"><sup>19</sup></a></li> <li><i>Cobitis laoensis</i> (<a href="/facts/Henri_%25C3%2589mile_Sauvage/hcILy4zC">Sauvage</a>, 1878)</li> <li><i>Cobitis laterimaculata</i> J. P. Yan & M. L. Zheng, 1984<a class="footnote-ref" id="fnref:20" href="#fn:20"><sup>20</sup></a></li> <li><i>Cobitis lebedevi</i> Vasil'eva & Vasil'ev, 1985</li> <li><i><a href="/facts/Cobitis_levantina/crc9ybye">Cobitis levantina</a></i> Krupp & Moubayed-Breil, 1992 (Orontes spined loach)</li> <li><i>Cobitis linea</i> (<a href="/facts/Johann_Jakob_Heckel/tjkqX0qp">Heckel</a>, 1847)</li> <li><i>Cobitis longibarba</i> (Y. F. Chen & Y. X. Chen, 2005)<a class="footnote-ref" id="fnref:21" href="#fn:21"><sup>21</sup></a></li> <li><i>Cobitis longicorpus</i> I. S. Kim, K. C. Choi & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1976<a class="footnote-ref" id="fnref:22" href="#fn:22"><sup>22</sup></a></li> <li><i>Cobitis lutheri</i> <a href="/facts/Carl_Hialmar_Rendahl/GeelLj2E">Rendahl</a> (de), 1935 (Luther's spined loach)<a class="footnote-ref" id="fnref:23" href="#fn:23"><sup>23</sup></a></li> <li><i>Cobitis macrostigma</i> Dabry de Thiersant, 1872</li> <li><i>Cobitis magnostriata</i> Nakajima, 2012<a class="footnote-ref" id="fnref:24" href="#fn:24"><sup>24</sup></a></li> <li><i><a href="/facts/Cobitis_maroccana/xz4dbTS4">Cobitis maroccana</a></i> <a href="/facts/Jacques_Pellegrin/AIMTMWcK">Pellegrin</a>, 1929</li> <li><i>Cobitis matsubarai</i> <a href="/facts/Yaichir%25C5%258D_Okada/XwQk0JsC">Okada</a> & Ikeda, 1939</li> <li><i><a href="/facts/Cobitis_megaspila/tSv3X8HN">Cobitis megaspila</a></i> <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1993</li> <li><i>Cobitis melanoleuca</i> <a href="/facts/John_Treadwell_Nichols/m3KgkrpI">Nichols</a>, 1925</li> <li><i><a href="/facts/Cobitis_meridionalis/kac8pyvm">Cobitis meridionalis</a></i> <a href="/facts/Stanko_Luka_Karaman/vIDSfl02">S. L. Karaman</a>, 1924</li> <li><i>Cobitis microcephala</i> Y. X. Chen & Y. F. Chen, 2011<a class="footnote-ref" id="fnref:25" href="#fn:25"><sup>25</sup></a></li> <li><i><a href="/facts/Cobitis_minamorii/zkpkb2bE">Cobitis minamorii</a></i> Nakajima, 2012<a class="footnote-ref" id="fnref:26" href="#fn:26"><sup>26</sup></a></li> <li><i>Cobitis minhi</i> S. V. Ngô, 2008</li> <li><i>Cobitis multifasciata</i> Wakiya & <a href="/facts/Tamezo_Mori/v5zL6r6y">T. Mori</a>, 1929<a class="footnote-ref" id="fnref:27" href="#fn:27"><sup>27</sup></a></li> <li><i>Cobitis multimaculata</i> Y. X. Chen & Y. F. Chen, 2011<a class="footnote-ref" id="fnref:28" href="#fn:28"><sup>28</sup></a></li> <li><i>Cobitis nalbanti</i> Vasil'eva, D. M. Kim, Vasil'ev, M. H. Ko & Y. J. Won, 2016 (Nalbant's spined loach)<a class="footnote-ref" id="fnref:29" href="#fn:29"><sup>29</sup></a></li> <li><i><a href="/facts/Cobitis_narentana/AnSz53CR">Cobitis narentana</a></i> <a href="/facts/Stanko_Luka_Karaman/vIDSfl02">S. L. Karaman</a>, 1928</li> <li><i><a href="/facts/Cobitis_ohridana/oIV5EsnU">Cobitis ohridana</a></i> <a href="/facts/Stanko_Luka_Karaman/vIDSfl02">S. L. Karaman</a>, 1928</li> <li><i>Cobitis pacifica</i> I. S. Kim, J. Y. Park & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1999<a class="footnote-ref" id="fnref:30" href="#fn:30"><sup>30</sup></a></li> <li><i><a href="/facts/Cobitis_paludica/kkSaEl6u">Cobitis paludica</a></i> (<a href="/facts/Don_Fernando_de_Buen_y_Lozano/Fq5QTaRD">F. de Buen</a>, 1930)</li> <li><i>Cobitis phrygica</i> <a href="/facts/Fahire_Battalgazi/Tt0oRIGd">Battalgazi</a>, 1944 (Acı spined loach)</li> <li><i>Cobitis pontica</i> Vasil'eva & Vasil'ev, 2006 (Pontic spined loach)</li> <li><i>Cobitis pumila</i> I. S. Kim & W. O. Lee, 1987 (Puan spined loach)<a class="footnote-ref" id="fnref:31" href="#fn:31"><sup>31</sup></a></li> <li><i><a href="/facts/Cobitis_puncticulata/yA03SFJg">Cobitis puncticulata</a></i> Erk' Akan, Ekmekçi & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1998 (Spotted spined loach)</li> <li><i><a href="/facts/Cobitis_punctilineata/aCoM59v3">Cobitis punctilineata</a></i> Economidis & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1996</li> <li><i>Cobitis rara</i> J. X. Chen, 1981</li> <li><i>Cobitis sakahoko</i> Nakajima & Suzawa, 2015<a class="footnote-ref" id="fnref:32" href="#fn:32"><sup>32</sup></a></li> <li><i>Cobitis saniae</i> Eagderi, Jouladeh-Roudbar, Jalili, Sayyadzadeh & Esmaeili, 2017<a class="footnote-ref" id="fnref:33" href="#fn:33"><sup>33</sup></a></li> <li><i>Cobitis satunini</i> Gladkov, 1935</li> <li><i>Cobitis shikokuensis</i> Suzawa, 2006</li> <li><i>Cobitis sibirica</i> Gladkov, 1935</li> <li><i>Cobitis simplicispina</i> <a href="/facts/B%25C3%25A9la_Hank%25C3%25B3/lwWpf4B5">Hankó</a> (hu), 1925 (Sakarya spined loach)</li> <li><i>Cobitis sinensis</i> <a href="/facts/Henri_%25C3%2589mile_Sauvage/hcILy4zC">Sauvage</a> & Dabry de Thiersant, 1874 (Siberian spined loach)</li> <li><i>Cobitis sipahilerae</i> Erk' Akan, Özdemir & Özeren, 2017<a class="footnote-ref" id="fnref:34" href="#fn:34"><sup>34</sup></a></li> <li><i>Cobitis splendens</i> Erk' Akan, Ekmekçi & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1998 (Splendid spined loach)</li> <li><i>Cobitis stenocauda</i> Y. X. Chen & Y. F. Chen, 2013<a class="footnote-ref" id="fnref:35" href="#fn:35"><sup>35</sup></a></li> <li><i><a href="/facts/Cobitis_stephanidisi/Y0Xcgl59">Cobitis stephanidisi</a></i> Economidis, 1992</li> <li><i><a href="/facts/Cobitis_striata/scJJPa9h">Cobitis striata</a></i> Ikeda, 1936<a class="footnote-ref" id="fnref:36" href="#fn:36"><sup>36</sup></a></li> <li><i><a href="/facts/Cobitis_strumicae/norojlSj">Cobitis strumicae</a></i> <a href="/facts/Stanko_Luka_Karaman/vIDSfl02">S. L. Karaman</a>, 1955 (Bulgarian spined loach)</li> <li><i><a href="/facts/Cobitis_taenia/fNsBgcny">Cobitis taenia</a></i> <a href="/facts/Carl_Linnaeus/gy5HWkHS">Linnaeus</a>, 1758 (Spined loach)</li> <li><i><a href="/facts/Cobitis_takatsuensis/MD4IDzxv">Cobitis takatsuensis</a></i> <a href="/facts/Nobuhiko_Mizuno/cWJEQQDt">Mizuno</a>, 1970</li> <li><i>Cobitis takenoi</i> Nakajima, 2016<a class="footnote-ref" id="fnref:37" href="#fn:37"><sup>37</sup></a></li> <li><i>Cobitis tanaitica</i> <a href="/facts/Mihai_C._B%25C4%2583cescu/0dMADKyp">Băcescu</a> & R. F. Mayer, 1969</li> <li><i>Cobitis taurica</i> Vasil'eva, Vasil'ev, Janko, Ráb & Rábová, 2005</li> <li><i>Cobitis tetralineata</i> I. S. Kim, J. Y. Park & <a href="/facts/Teodor_T._Nalbant/Lu41duvD">Nalbant</a>, 1999</li> <li><i><a href="/facts/Cobitis_trichonica/wzJCBhAG">Cobitis trichonica</a></i> Stephanidis, 1974</li> <li><i><a href="/facts/Cobitis_turcica/u4AErBGH">Cobitis turcica</a></i> <a href="/facts/B%25C3%25A9la_Hank%25C3%25B3/lwWpf4B5">Hankó</a> (hu), 1925 (Turkish spined loach)</li> <li><i><a href="/facts/Cobitis_vardarensis/abqLrgvD">Cobitis vardarensis</a></i> <a href="/facts/Stanko_Luka_Karaman/vIDSfl02">S. L. Karaman</a>, 1928 (Vardar spined loach)</li> <li><i><a href="/facts/Cobitis_vettonica/5fH4Ncrd">Cobitis vettonica</a></i> Doadrio & Perdices, 1997</li> <li><i>Cobitis xinjiangensis</i> (Y. F. Chen & Y. X. Chen, 2005)<a class="footnote-ref" id="fnref:38" href="#fn:38"><sup>38</sup></a></li> <li><i>Cobitis ylengensis</i> S. V. Ngô, 2003</li> <li><i>Cobitis yongdokensis</i> (I. S. Kim & J. Y. Park, 1997)<a class="footnote-ref" id="fnref:39" href="#fn:39"><sup>39</sup></a></li> <li><i><a href="/facts/Cobitis_zanandreai/byvxW9yW">Cobitis zanandreai</a></i> Cavicchioli, 1965</li> <li><i>Cobitis zhejiangensis</i> Y. M. Son & S. P. He, 2005</li></ul> <h2 id="references">References</h2> <ol> <li id="fn:1"><p>Froese, Rainer; Pauly, Daniel (eds.). "Species in genus Cobitis". FishBase. June 2017 version. <a href="/wiki/Rainer_Froese" target="_blank">/wiki/Rainer_Froese</a> <a href="#fnref:1" class="footnote-back-ref">↩</a></p></li> <li id="fn:2"><p>Perdices, A., Bohlen, J. & Doadrio, I. (2008): The molecular diversity of adriatic spined loaches (Teleostei, Cobitidae). Molecular Phylogenetics and Evolution, 46 (1): 382–390. <a href="http://www.sciencedirect.com/science/article/pii/S1055790307001650" target="_blank">http://www.sciencedirect.com/science/article/pii/S1055790307001650</a> <a href="#fnref:2" class="footnote-back-ref">↩</a></p></li> <li id="fn:3"><p>Vasil'eva, E.D. & Vasil'ev, V.P. (2012): Cobitis amphilekta sp. nova, a New Species of Spined Loaches (Cobitidae, Cypriniformes) from the Caspian Sea Basin. Journal of Ichthyology, 52 (3): 200–206. <a href="https://link.springer.com/article/10.1134/S0032945212020154" target="_blank">https://link.springer.com/article/10.1134/S0032945212020154</a> <a href="#fnref:3" class="footnote-back-ref">↩</a></p></li> <li id="fn:4"><p>Chen, Y.-X., Chen, Y.-F. & He, D.-K. (2013): A new species of spined loach (Osteichthyes, Cobitidae) from the Pearl River, Guangxi of China. Acta Zootaxonomica Sinica, 38 (2): 377-387. <a href="#fnref:4" class="footnote-back-ref">↩</a></p></li> <li id="fn:5"><p>Mousavi-Sabet, H., Vatandoust, S., Esmaeili, H.R., Geiger, M.F. & Freyhof, J. (2015): Cobitis avicennae, a new species of spined loach from the Tigris River drainage (Teleostei: Cobitidae). Zootaxa, 3914 (5): 558–568. <a href="http://www.mapress.com/zootaxa/2015/f/z03914p568f.pdf" target="_blank">http://www.mapress.com/zootaxa/2015/f/z03914p568f.pdf</a> <a href="#fnref:5" class="footnote-back-ref">↩</a></p></li> <li id="fn:6"><p>Chen, Y., Sui, X., Liang, N. & Chen, Y. (2015): Two new species of the genus Cobitis Linnaeus (Teleostei: Cobitidae) from southern China. Chinese Journal of Oceanology and Limnology, 34 (3): 517–525. <a href="https://link.springer.com/article/10.1007/s00343-016-4277-0" target="_blank">https://link.springer.com/article/10.1007/s00343-016-4277-0</a> <a href="#fnref:6" class="footnote-back-ref">↩</a></p></li> <li id="fn:7"><p>Perdices, A., Bohlen, J., Šlechtová, V. & Doadrio, I. (2016): Molecular Evidence for Multiple Origins of the European Spined Loaches (Teleostei, Cobitidae). PLoS ONE, 11 (1): e0144628. <a href="http://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0144628&type=printable" target="_blank">http://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0144628&type=printable</a> <a href="#fnref:7" class="footnote-back-ref">↩</a></p></li> <li id="fn:8"><p>Chen, Y. & Chen, Y. (2013): Three new species of cobitid fish (Teleostei, Cobitidae) from the River Xinjiang and the River Le’anjiang, tributaries of Lake Poyang of China, with remarks on their classification. Folia Zoologica, 62 (2): 83-95. <a href="http://www.ivb.cz/folia_zoologica/archive/62_83-95.pdf" target="_blank">http://www.ivb.cz/folia_zoologica/archive/62_83-95.pdf</a> <a href="#fnref:8" class="footnote-back-ref">↩</a></p></li> <li id="fn:9"><p>Erkakan, F. & Özdemir, F. (2014): The first new cave fish species, Cobitis damlae (Teleostei: Cobitidae) from Turkey. Hacettepe Journal of Biology and Chemistry, 42 (2): 275–279. <a href="http://www.hjbc.hacettepe.edu.tr/site/assets/files/1452/hjbc_42_2_275-279.pdf" target="_blank">http://www.hjbc.hacettepe.edu.tr/site/assets/files/1452/hjbc_42_2_275-279.pdf</a> <a href="#fnref:9" class="footnote-back-ref">↩</a></p></li> <li id="fn:10"><p>Perdices, A., Bohlen, J., Šlechtová, V. & Doadrio, I. (2016): Molecular Evidence for Multiple Origins of the European Spined Loaches (Teleostei, Cobitidae). PLoS ONE, 11 (1): e0144628. <a href="http://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0144628&type=printable" target="_blank">http://journals.plos.org/plosone/article/file?id=10.1371/journal.pone.0144628&type=printable</a> <a href="#fnref:10" class="footnote-back-ref">↩</a></p></li> <li id="fn:11"><p>Erkakan, F., Özdemir, F. & Özeren, S.C. (2017): Two new species of the genus Cobitis Linnaeus (Teleostei: Cobitidae) from Turkey. FishTaxa, 2 (2): 82-89. <a href="http://fishtaxa.com/index.php/ft/article/view/2-2-4/87" target="_blank">http://fishtaxa.com/index.php/ft/article/view/2-2-4/87</a> <a href="#fnref:11" class="footnote-back-ref">↩</a></p></li> <li id="fn:12"><p>Mousavi-Sabet, H., Vasil'eva, E.D., Vatandoust, S. & Vasil'ev, V.P. (2011): Cobitis faridpaki sp. nova — a New Spined Loach Species (Cobitidae) from the Southern Caspian Sea Basin (Iran). 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