The most intense severe weather activity was expected to come across the southern Great Plains, specifically Central Oklahoma, during the afternoon hours on that Monday. As such, the Storm Prediction Center issued a moderate risk of severe thunderstorms during the early morning hours of May 20 from southeastern Missouri to north-central Texas. The degree of wind shear, moisture, and instability within the warm sector favored the development of supercells. Very large hail and tornadoes were both expected with these supercells, with the possibility of a few strong tornadoes. The Storm Prediction Center issued a tornado watch at 1:10 p.m. Central Daylight Time (CDT) early that afternoon for the eastern two-thirds of Oklahoma, northwestern Arkansas, and portions of north-central Texas. Given the atmospheric parameters thought to be in place at the time, the Storm Prediction Center inadvertently underestimated the threat of tornadic activity that afternoon; the probability table for the tornado watch – the 191st severe weather watch issued by the guidance center in 2013 – indicated a 40% (or "moderate") probability of two or more tornadoes and a 20% (or "low") probability of one or more tornadoes reaching between EF2 and EF5 intensity within the watch area. The watch did, however, say that "one or two strong tornadoes" were possible.
The thunderstorm that eventually produced the tornado developed less than one hour after the tornado watch was issued, around 2:00 p.m. CDT, across northern Grady County. Its rapid intensification resulted in the National Weather Service Weather Forecast Office in Norman issuing a severe thunderstorm warning for northern Grady, northwestern Cleveland, northern McClain, and southwestern Oklahoma Counties (including southwestern portions of the immediate Oklahoma City area) at 2:12 p.m. CDT. The thunderstorm quickly attained supercell characteristics, with rotation at the mid-levels of the storm's cloud structure becoming apparent even before it was officially classified as severe, due to the sufficient amounts of wind shear present over central Oklahoma.
At 2:40 p.m. CDT, as rotation in the supercell was increasing at the cloud base, a tornado warning was issued for far northeastern Grady, western Cleveland, northern McClain, and southern Oklahoma Counties, as the storm approached the Oklahoma City metropolitan area. The rapid formation of the storm may have been associated with a descending reflectivity core and interplay of complex cell mergers.
The tornado touched down at 2:56 p.m. CDT in northwestern McClain County, near Long Drive to the southwest of the North Country Club Road and SH-37 intersection, roughly 4.4 miles (7.1 km) northwest of downtown Newcastle. Initially a narrow rope-shaped tornado, it first caused EF1 damage to a home and some trees before crossing SH-37 and rapidly intensifying as it struck a semi-rural subdivision, where several homes were destroyed, two of which were leveled at EF4 intensity. At this point, the tornado began to grow rapidly in size over the course of 2 minutes; the rope-like funnel started to swell into a large stovepipe, before maturing further into a barrel shape. A debris ball grew around the funnel itself as it finally evolved into a large, wedge-shaped structure. Slight ground scouring began in this area, and several homes in a subdivision further to the northeast sustained EF3 damage. By 3:01 p.m. CDT, the National Weather Service issued a second, more strongly worded warning for the area: in a severe weather statement updating the existing tornado warning, the Norman forecast office declared a tornado emergency for southern Oklahoma City and Moore as various storm spotters confirmed that the large, violent tornado was approaching the area. The Twin Lakes dual-polarization NEXRAD radar (near Lake Stanley Draper) detected a tornado debris signature one mile in diameter within the accompanying hook echo, as the tornado crossed into portions of south Oklahoma City in northern Cleveland County. The supercell also produced straight-line winds of 80 miles per hour (130 km/h) for at least one mile surrounding the tornadic circulation.
The tornado maintained its intensity as it struck the Orr Family Farm and the Celestial Acres horse training area, where up to 100 horses were reported killed, some being tossed into and tangled in downed power lines or thrown on top of nearby buildings; several horses that survived the tornado suffered severe injuries, with some being impaled by tree limbs or boards. Every building at Celestial Acres was either leveled or swept away at EF4 intensity, the ground on the property was scoured to bare soil, debris from structures was granulated, and vehicles were thrown and stripped down to their frames. Surveyors noted that based on the contextual damage, the tornado was likely at EF5 intensity in this area, though the construction quality of the affected buildings only permitted an EF4 rating. A 10-ton propane tank on the Orr Farm property was picked up and thrown more than a half-mile through the air by the tornado, and a strip mall near the farm was completely leveled as well. The tornado continued east, heavily scouring an open grassy field before impacting Briarwood Elementary School, which was completely destroyed. The NWS originally rated this damage EF5, but further evaluation and a 2014 study published by the American Meteorological Society revealed evidence of poor construction at the school, and the rating was downgraded to EF4. Remarkably, no fatalities occurred at the school. Two 12,000-gallon water tanks that were also swept off of the Orr Family Farm grounds were thrown into this area; the roof of Briarwood Elementary was struck by one of them − potentially aiding in compromising the building's structural integrity as it bent the steel girders that held up the roof − shortly before the main vortex struck the building, while the other fell onto and destroyed a home a few blocks east of the school. Past Briarwood Elementary, the tornado intensified even further and entered densely populated areas just west of Moore, including the Westmoor subdivision where many well-built, anchor-bolted brick homes were flattened at EF4 intensity, and two were swept clean from their foundations, with damage at those two homes rated EF5.
The tornado continued generally northeast at EF4 strength, completely debarking trees and leveling entire neighborhoods as it moved into the western city limits of Moore. Many homes were flattened in neighborhoods to the east of South Santa Fe Avenue. The tornado was initially rated EF5 in this area as only the bare slab of an anchor-bolted home remained. However, it was later downgraded to an EF4 as an inspection revealed the home's anchor bolts were missing their nuts and washers. The tornado then destroyed Plaza Towers Elementary School at EF4 intensity where seven children were killed when a cinder block wall collapsed on top of them. More than a dozen homes in a subdivision just to the south of Plaza Towers Elementary were completely swept away, all revealed to have been nailed rather than bolted to their foundations. Damage to this subdivision was subsequently rated EF4, though the tornado was likely violent as lawns were completely scoured down to bare soil. Entire blocks of homes were flattened, trees were completely debarked and denuded, vehicles were thrown and mangled, and the ground was severely scoured in other residential areas nearby. The damage also earned an EF4 rating in these areas. Most of the fatalities from the tornado occurred in the Plaza Towers neighborhood of Moore. In one of these houses (a block away from Plaza Towers Elementary), a woman was killed as she tried to seek shelter in a closet.
Further to the northeast, at least a dozen cars were piled up against the front entrance of the Moore Medical Center, which sustained EF4 damage. One car was lofted and thrown onto the roof. Many homes in neighborhoods near the medical center were completely destroyed, including a row of four well-built brick homes with anchor bolts that were swept away, with damage to those four homes rated EF5. An open field directly behind this row of homes was deeply scoured, with only bare soil and clumps of dirt remaining. A nearby manhole cover was removed, and multiple vehicles were mangled beyond recognition and caked in mud in this area as well. The nearby Warren Theater was spared a direct hit, but still sustained considerable damage to its exterior. A bowling alley in the area was leveled, and a 7-Eleven which was on the northern part of the damage path, was completely flattened with four people killed inside (including a three-month-old infant). The nearby Moore Cemetery was heavily damaged as well.
The tornado briefly weakened and caused EF3 damage to some other businesses near I-35/US 77, before crossing the interstate and mangling several vehicles in the process. The tornado regained EF4 intensity on the other side of the Interstate as it tore through several neighborhoods and destroyed numerous additional homes (though the EF4 damage swath was narrower at this point). A large grassy field between two subdivisions in this area was scoured to bare soil, with wind-rowed structural debris and several mangled vehicles strewn to the east. One brick home on Hunters Glen Court sustained EF5 damage, with only the slab foundation and anchor bolts remaining. Very little structural debris or house contents was recovered from that residence, and the small number of debris that remained was wind-rowed well away from the site. Two vehicles were also lofted from the residence, one of which was thrown over 100 yards (91 m).
The tornado continued through Moore's eastern neighborhoods. Highland East Jr. High's main building was spared, but the separate gymnasium building was completely destroyed, and a set of lockers from the structure was lofted and thrown a considerable distance into a nearby neighborhood. The Moore Public Schools administration building, a converted former hospital located a few blocks to the east, was also struck and destroyed. The tornado was noticeably narrower at this point, but was still causing a continuous swath of EF4 damage to numerous homes as it moved through multiple subdivisions. A well-built, anchor-bolted home at the corner of Heatherwood Drive and Southeast 5th Street was reduced to a bare slab, sustaining EF5 damage. A large, well-bolted-down home at the end of a private drive off of Southeast 4th Street (SH-37) was also swept cleanly away at EF5 intensity. Debris was scattered well away from the site, a vehicle was thrown over 100 yards (91 m), and wind-rowing was again noted at that location.
Further east, the tornado weakened to EF3 strength as it exited Moore, destroying six industrial buildings and damaging two others. A final small area of EF4 damage was noted nearby as two homes and a concrete building were leveled. The tornado then began to rapidly narrow and weaken, snapping and uprooting several trees and causing EF2 damage to a farm just east of Moore, where the house lost its roof and an outbuilding was destroyed. A pickup truck slid 200 feet (61 m) away from the farm into a field while remaining upright. At 3:35 p.m. CDT, the tornado dissipated at a nearby tree line about one-third mile (0.54 km) east of South Air Depot Boulevard. The tornado traveled a total of 13.85 miles (22.3 km) over 39 minutes,: 13 moving at an average rate of 21 miles per hour (34 km/h).: 4 It reached a maximum width of 1,900 yards (1,737 m).: 13
The violent updrafts in the supercell that produced the tornado lofted debris from homes and other buildings in Moore at least 10 miles (16 km) outside of the tornado's outer circulation, with reports of various objects and personal possessions being found as far away as Midwest City.
At Plaza Towers, the second school to be hit, parents had been allowed to collect their children early in preparation for the oncoming storm. Therefore, by the time the tornado struck only about 75 students and teachers were in the building. Many students and teachers took shelter in bathrooms and closets, but in a newer addition to the building which housed the school's second and third grade classrooms, seven fatalities occurred. Third grade teacher Jennifer Doan was taking shelter with eleven of her students in a hallway when the tornado struck. Doan and her students were trapped when the walls of the corridor collapsed on top of them. Doan, who was two months pregnant with her third child at the time, suffered severe injuries to her back, but did not lose the baby. She and five of her students were pulled out, with the children suffering only minor to moderate injuries. Another six of her students along with a student from another third-grade class died.
The Moore Medical Center was heavily damaged, but no one present in the building was injured by the tornado. The center's staff had to relocate 30 patients to a hospital in Norman and another hospital. Part of I-35 was shut down due to debris that had been thrown onto the freeway. On May 21, Moore still did not have running water. There were more than 61,500 power outages related to the tornado. More than 100 people were rescued from areas that sustained significant damage from the tornado.
In addition to erroneous reports concerning the status of Briarwood and Plaza Towers students, various reports regarding fatality and injury counts as well as persons unaccounted for were circulated by traditional and social media outlets in the immediate hours after the tornado struck, which could not immediately be confirmed or refuted due to communications disruptions in the affected areas or were the result of incorrect counting due to miscommunication of estimates. Within the first two days, it was reported that between 237 and 240 people had been injured, with the tally later increasing to over 350. The final count of injuries was later adjusted downward to 212.
The Moore City Council proposed a measure making twelve changes to its residential building codes, include requiring that new home construction in the city include hurricane clips or framing anchors, continuous plywood bracing and wind-resistant garage doors in order for homes to withstand winds up to 135 miles per hour (217 km/h) (equivalent to a high-end EF2 tornado). When the measure was passed in a unanimous vote held on March 17, 2014, Moore became the first city in the United States to adopt a building code addressing the effects of tornadoes on homes, which exceed the national standards set by the National Association of Home Builders.
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