By 2001, Slash's Snakepit had disbanded for the second time. Slash began working with the Black Crowes drummer Steve Gorman and an unnamed bassist on a new project: writing the music for what would become "Fall to Pieces". McKagan reformed Loaded, previously his band for the tour in support of Beautiful Disease, with Geoff Reading. McKagan also added both Mike Squires and Jeff Rouse to the lineup. Following a tour of Japan in 2002, former Zilch, Wasted Youth, Electric Love Hogs, and Dave Navarro guitarist Dave Kushner joined Loaded in place of Mike Squires.
While Shaefer's audition was unsuccessful, the quartet continued auditioning for a lead singer. VH1 filmed the recruitment process, which was referred to as the temporary name "The Project". The resulting documentary was aired as VH1 Inside Out: The Rise of Velvet Revolver. A number of lead singers auditioned for the band, including Stephen Shareaux of Kik Tracee, Steve Ludwin of Carrie and Little Hell, Todd Kerns, formerly of Age of Electric, Sebastian Bach, formerly of Skid Row, Shawn Albro of U.P.O., Travis Meeks of Days of the New, and Ian Thornley of Big Wreck. Myles Kennedy, formerly of the Mayfield Four, declined an invitation from Sorum to audition. Ian Astbury of the Cult and Mike Patton of Faith No More also declined audition offers. The band were also interested in auditioning Stone Temple Pilots singer Scott Weiland, who had become friends with McKagan after attending the same gym. Weiland once played on the same bill as Kushner, and was in rehab at the same time as Sorum. Weiland was sent two discs of material, and felt that the first disc "sounded like Bad Company gone wrong." When he was sent the second disc, Weiland was more positive, comparing it to Core-era Stone Temple Pilots, though he turned them down because Stone Temple Pilots were still together.
When Stone Temple Pilots disbanded in 2003, the band sent Weiland new music, which he took into his studio and added vocals. This music eventually became the song "Set Me Free". Weiland was still unsure whether or not he wanted to join, despite delivering the music to the band himself and performing at an industry showcase at Mates. They recorded two songs with producer Nick Raskulinecz, a recorded version of "Set Me Free" and a cover of Pink Floyd's "Money", for the soundtracks to the movies Hulk and The Italian Job, respectively. Weiland joined the band soon after. "Set Me Free" managed to peak at number 17 on the Mainstream Rock Chart without any radio promotion or a record label.
Prior to the recording of their debut album, Weiland took material that the band had previously written to his studio, Lavish, in Toluca Lake. With engineer Doug Grean, Weiland rearranged the music to fit his vocals, eventually coming out with the songs "Big Machine" and "Dirty Little Thing". The band worked on new material for songs such as "You Got No Right", "Slither", "Sucker Train Blues", and "Do It for the Kids", among others. During this time Weiland was arrested at the parking lot of his studio for drug possession. Upon release from jail, he wrote lyrics to material he was given previously, writing the lyrics to the song "Fall to Pieces". Velvet Revolver soon began recording their debut album. Initially, they recorded "Slither" with producer Bob Ezrin at Henson Studios, but were dissatisfied with the result. After recording "Headspace" with Josh Abraham, the band liked the track enough to do the rest of the album with him.
Critically, the album was generally well received. Despite being praised for its hedonism and maturity, critics noted a disconnection between "singer and band". Velvet Revolver won the Kerrang! Award for Best International Newcomer in 2004, and the following year they won the Best Hard Rock Performance Grammy Award for "Slither". They were also nominated for Rock Artist of the Year at the Billboard Music Awards while "Fall to Pieces" was nominated for a Song of the Year/Rock Radio Radio Music Award. They recorded a new song entitled "Come On, Come In" for the movie Fantastic Four in 2005, which peaked at number 14 on the Mainstream Rock Chart. "Fall to Pieces" then re-entered the charts, peaking at number twenty-five on the Adult Top 40 the same year.
Following the completion of the album, Velvet Revolver performed for and inducted Van Halen into the Rock and Roll Hall of Fame, with Weiland and Slash speaking on the band's behalf, on March 12, 2007. The band played a medley of "Ain't Talkin' 'bout Love" and "Runaround". Shows in South America with Aerosmith followed in April. They released the EP Melody and the Tyranny on June 1 to serve as a precursor to the release of their new album, which featured two songs from Libertad, a cover of Talking Heads song "Psycho Killer" and a video documentary about the making of Libertad as well as a live video of the band performing "Do It for the Kids".
On the UK tour, the band members never spoke with Weiland, with the exception of a few arguments around the stage. Tensions came to a head during Velvet Revolver's Glasgow show on March 20, 2008, where Weiland announced to the crowd that it was the band's last tour, unaware that the other band members were already planning on firing him. After Sorum posted a message about the show on his website, Weiland issued a statement through Blabbermouth.net in response, saying he "made many attempts to remain cordial with the members of [Velvet Revolver], but mainly, the likes of [Sorum]" and that "[the band] were a gang. But ego and jealousy can get the better of anyone." Slash later stated that it would not be Velvet Revolver's last tour. Weiland's departure was announced on April 1. Weiland also departed the cover band Camp Freddy, which also featured Sorum, and reunited with Stone Temple Pilots, before being fired in 2013.
After Weiland's departure, the band began recording and searching for a new singer. The search was sporadic with the band spending some time auditioning singers, then turning into solo projects, returning to the band, then abandoning it again. Several names were rumored to be auditioning for the band through the years. Myles Kennedy (Alter Bridge) was strongly rumored due to his collaborations with Slash; Lenny Kravitz, Chester Bennington of Linkin Park, Steve Isaacs formerly of Skycycle and the Panic Channel, Royston Langdon of Spacehog, Donovan Leitch of Camp Freddy, Ours singer Jimmy Gnecco and Scars on Broadway guitarist Franky Perez, as well as previous auditionee Sebastian Bach. Ex-Slash's Snakepit lead vocalist Rod Jackson was rumoured to audition, but Slash stated that he would not collaborate with Jackson again, citing his poor work ethic and unreliability. Kushner later revealed that Perez was officially hired by the band as vocalist for a brief time in 2008.
Velvet Revolver reunited for a one-off performance with Scott Weiland at a benefit concert for the late John O'Brien, on January 12, 2012. Following a benefit show for the Road Recovery in 2011 with the other Velvet Revolver members, each one agreed to a one-off reunion before Kushner invited Weiland, who also agreed. Kushner also stated it was then unknown what Velvet Revolver's plans were for the future after the reunion show; "I know everyone's got other commitments, but I think everyone's like, 'Let's get this thing done and get through this and then we'll see.'"
In April 2012, Weiland remarked that he would like to reunite permanently with Velvet Revolver, saying that "if Maynard James Keenan can do it with A Perfect Circle and Tool, then there's no reason why I shouldn't go and do it with both bands". Further in May in an interview with ABC Radio Weiland said that he had reunited with the band permanently for a tour and an album, which was denied a few days later by Slash in an interview with 93X.
On December 3, 2015, Weiland was found dead on his tour bus around 9 pm, one day before he was to perform in Minnesota with his band the Wildabouts, which ended the possible reunion. Both Slash and McKagan rejoined Guns N' Roses in 2016.
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