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Fall webworm
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<h2 id="description">Description</h2> <p>The adult fall webworm has a wingspan of approximately 30 mm and is generally white-colored, although some individuals may have dark-colored marks.<a class="footnote-ref" id="fnref:2" href="#fn:2"><sup>2</sup></a> </p> <h2 id="distribution">Distribution</h2> <p>The moth is native to <a href="/facts/North_America/djNjCQIl">North America</a>, ranging from Canada to Mexico and has been <a href="/facts/Introduced_species/s0aHefyp">introduced</a> into other continents.<a class="footnote-ref" id="fnref:3" href="#fn:3"><sup>3</sup></a> Introduced to what was formerly Yugoslavia in the 1940s (firstly recorded in 1949<a class="footnote-ref" id="fnref:4" href="#fn:4"><sup>4</sup></a>), it now has occupied probably its entire range in Europe from France to the Caspian Sea in the east as well as penetrated into <a href="/facts/Central_Asia/rB8k0CyT">Central Asia</a>: <a href="/facts/Turkmenistan/YqJ4NvjU">Turkmenistan</a> (from 1990 to 1993), <a href="/facts/Uzbekistan/o8wfw2xR">Uzbekistan</a> (Fergana valley from 1996 to 1997), <a href="/facts/Kyrgyzstan/MSNdCp8u">Kyrgyzstan</a>, and southeastern <a href="/facts/Kazakhstan/Vfs8NI6e">Kazakhstan</a>. It was also introduced into Japan in 1945 and has adjusted its number of generations per year since its arrival.<a class="footnote-ref" id="fnref:5" href="#fn:5"><sup>5</sup></a> It spread into China, southern Mongolia, Korea and southern <a href="/facts/Primorsky_Krai/a0gC2rCJ">Primorsky Krai</a> of <a href="/facts/Russia/jfkCwyxn">Russia</a> so that now it is considered <a href="/facts/Holarctic/uNgFyNx9">holarctic</a> in distribution.<a class="footnote-ref" id="fnref:6" href="#fn:6"><sup>6</sup></a> </p> <h2 id="life-cycle">Life cycle</h2> <p>One generation per year emerges in the northern part of <a href="/facts/North_America/djNjCQIl">North America</a>, with larvae appearing in late summer through early fall. South of an approximate <a href="/facts/40th_parallel_north/LLETD40F">latitude of 40°N</a> there are two or more generations annually, with webs appearing progressively earlier further south.<a class="footnote-ref" id="fnref:7" href="#fn:7"><sup>7</sup></a> The second generation is most damaging in southeast Pennsylvania and Delaware and emerges between 791-1540 <a href="/facts/Growing_degree-day/24XssIU8">growing degree-days</a>.<a class="footnote-ref" id="fnref:8" href="#fn:8"><sup>8</sup></a> </p> <h3>Eggs</h3> <p>The adult moth lays her eggs on the underside of leaves in 'hair'-covered clusters of a few hundred.<a class="footnote-ref" id="fnref:9" href="#fn:9"><sup>9</sup></a> Eggs hatch in about a week.<a class="footnote-ref" id="fnref:10" href="#fn:10"><sup>10</sup></a> </p> <h3>Larva</h3> <p>The <a href="/facts/Caterpillar/PMgzj7F7">caterpillars</a> are highly variable in coloration, ranging from a pale yellow to dark grey, with yellow spots and long and short bristles.<a class="footnote-ref" id="fnref:11" href="#fn:11"><sup>11</sup></a> There are two cream stripes along the sides. The two races—one more common in the north, the other in the south—differ in head capsule coloration.<a class="footnote-ref" id="fnref:12" href="#fn:12"><sup>12</sup></a> The maximum length of larvae is 35 mm. Webs are progressively enlarged and much messier looking than those of <a href="/facts/Malacosoma/fLJyHvlr">tent caterpillars</a> (which occur only in spring and have shorter hairs and very little yellow on their bodies); also, webs from the fall webworm are concentrated to the tips of the branches, whereas the tent caterpillar webs are largely found in the unions. Larvae feed inside the tents until the late <a href="/facts/Instar/KRIfCFIn">instars</a>. Very young larvae feed only on the upper surfaces of leaves; later, they consume whole leaves. The larval stage lasts about four to six weeks.<a class="footnote-ref" id="fnref:13" href="#fn:13"><sup>13</sup></a> Larvae are known to wiggle vigorously at periodic intervals in synchrony. How they synchronize these movements especially when distributed over a wide area has not been established.<a class="footnote-ref" id="fnref:14" href="#fn:14"><sup>14</sup></a> </p> <h3>Pupa</h3> <p>The pupa stage overwinters in the bark and leaf litter at the base of the trees. It is dark brown and about 10 mm long. The thin brown cocoon is made of silk with bits of detritus interwoven.<a class="footnote-ref" id="fnref:15" href="#fn:15"><sup>15</sup></a> </p> <h3>Adult</h3> <p>The adult is mostly white in the North America regions, but in the south it may be marked with black or brown spots on the forewings.<a class="footnote-ref" id="fnref:16" href="#fn:16"><sup>16</sup></a><a class="footnote-ref" id="fnref:17" href="#fn:17"><sup>17</sup></a> It is quite 'hairy' and the front legs have bright yellow or orange patches. The underwings will have less marking than the forewings, and the abdomen often has a sprinkling of brown hairs. It has a wingspan with a range of 35–42 mm. </p> <h2 id="food-plants">Food plants</h2> <p>The fall webworm feeds on just about any type of <a href="/facts/Deciduous/7jQHVtdd">deciduous</a> tree, where leaves are chewed; branches or the entire tree may become defoliated. Worldwide, it has been recorded from 636 species,<a class="footnote-ref" id="fnref:18" href="#fn:18"><sup>18</sup></a> and it is considered to be among the most polyphagous of insects. In the eastern U.S., <a href="/facts/Pecan/JnaYpv9I">pecan</a> trees (<i>Carya illinoinensis</i>), <a href="/facts/Juglans_nigra/wX0gYu1S">black walnut</a>, American elm (<i><a href="/facts/Ulmus_americana/Bjj7ercR">Ulmus americana</a></i> L.), hickory, fruit trees, and some maples are preferred hosts; in some areas persimmon and sweetgum are also readily eaten. In the west, alder, willow (<a href="/facts/Salix_sepulcralis_group/g90uyGm0"><i>Salix</i> spp.</a>), cottonwood and fruit trees are commonly used. Additional host plant examples include madrone (<i><a href="/facts/Arbutus_menziesii/r3Lo7arK">Arbutus menziesii</a></i> Pursh), mulberry, ailanthus, American sycamore (<i><a href="/facts/Platanus_occidentalis/aFtF4QPl">Platanus occidentalis</a></i> L.), and Asian white birch (<i><a href="/facts/Betula_platyphylla/I1VWxcF1">Betula platyphylla</a></i>).<a class="footnote-ref" id="fnref:19" href="#fn:19"><sup>19</sup></a><a class="footnote-ref" id="fnref:20" href="#fn:20"><sup>20</sup></a><a class="footnote-ref" id="fnref:21" href="#fn:21"><sup>21</sup></a> <a class="footnote-ref" id="fnref:22" href="#fn:22"><sup>22</sup></a> </p><p>Host plant selection is dependent on factors like the plant's degree of sun exposure, age, environmental stress undergone, toughness, and nutritional quality.<a class="footnote-ref" id="fnref:23" href="#fn:23"><sup>23</sup></a><a class="footnote-ref" id="fnref:24" href="#fn:24"><sup>24</sup></a> For example, for an insect that needs energy for processes like dispersal or <a href="/facts/Diapause/y4fPYEjC">diapause</a>, consuming plants that provide a lot of carbohydrates could be beneficial; for a female insect that is producing eggs, consuming plants that provide a lot of protein could be beneficial.<a class="footnote-ref" id="fnref:25" href="#fn:25"><sup>25</sup></a> </p> <h2 id="behavior">Behavior</h2> <p>The fall webworm is a gregarious insect.<a class="footnote-ref" id="fnref:26" href="#fn:26"><sup>26</sup></a> Groups of larvae live in self-created large webs that are found on tree branches.<a class="footnote-ref" id="fnref:27" href="#fn:27"><sup>27</sup></a><a class="footnote-ref" id="fnref:28" href="#fn:28"><sup>28</sup></a> These webs allow for the finding of mates, temperature regulation, increased growth rate, and protection from predators, but also result in higher rates of infections and predation.<a class="footnote-ref" id="fnref:29" href="#fn:29"><sup>29</sup></a><a class="footnote-ref" id="fnref:30" href="#fn:30"><sup>30</sup></a> </p><p>The caterpillars of the fall webworm have several strategies to defend against threats. Some examples of protective behavior or defense are shaking and jerking together, repellant scent, and irritants on hairs or spines.<a class="footnote-ref" id="fnref:31" href="#fn:31"><sup>31</sup></a><a class="footnote-ref" id="fnref:32" href="#fn:32"><sup>32</sup></a> </p><p>The fall webworm exhibits a form of parental care, where the female will try to protect the eggs after oviposition by covering her newly laid eggs with her abdomen hairs.<a class="footnote-ref" id="fnref:33" href="#fn:33"><sup>33</sup></a> </p> <h2 id="reproduction">Reproduction</h2> <h3>Mating limitations</h3> <p>Male fall webworms are typically only available for mating for about 30–60 minutes per day.<a class="footnote-ref" id="fnref:34" href="#fn:34"><sup>34</sup></a> </p> <h3>Sex pheromone</h3> <p>The <a href="/facts/Sex_pheromone/fcAmVKxG">sex pheromone</a> consists of the components (9Z,12Z)- 9,12-octadecadienal (I), (9Z,12Z,15Z)-9,12,15-octadecatrienal (II), cis-9,10-epoxy-(3Z,6Z)-3,6-henicosa- diene (III), and cis-9,10-epoxy-(3Z,6Z)-1,3,6-henicosatriene (IV).<a class="footnote-ref" id="fnref:35" href="#fn:35"><sup>35</sup></a> </p> <h2 id="physiology">Physiology</h2> <h3>Thermoregulation</h3> <p>Fall webworms experience <a href="/facts/Behavioural_thermoregulation/V692hIjw">behavioral thermoregulation</a>.<a class="footnote-ref" id="fnref:36" href="#fn:36"><sup>36</sup></a><a class="footnote-ref" id="fnref:37" href="#fn:37"><sup>37</sup></a> The fall webworms' self-created web (which is where the fall webworms live) is able to trap heat.<a class="footnote-ref" id="fnref:38" href="#fn:38"><sup>38</sup></a><a class="footnote-ref" id="fnref:39" href="#fn:39"><sup>39</sup></a> Due to this, the fall webworm (which is an <a href="/facts/Ectotherm/vdjXLuQr">ectotherm</a>) is able to maintain a warm temperature of about 40-50 °C, which allows the larvae to grow and develop faster.<a class="footnote-ref" id="fnref:40" href="#fn:40"><sup>40</sup></a><a class="footnote-ref" id="fnref:41" href="#fn:41"><sup>41</sup></a> Inside the web, there is a temperature difference because the central part tends to have a higher temperature, while the back part tends to have a lower temperature.<a class="footnote-ref" id="fnref:42" href="#fn:42"><sup>42</sup></a> However, a web's heat trapping process is not always stable; wind can disturb the heat trapping.<a class="footnote-ref" id="fnref:43" href="#fn:43"><sup>43</sup></a> </p> <h3>Intestinal tract</h3> <p>There are several components to the fall webworm gut, such as the foregut and the midgut. The foregut and midgut of the fall webworm are alkaline, and Johnson and Felon found that the midgut pH ranged from 8.7 to 11.4.<a class="footnote-ref" id="fnref:44" href="#fn:44"><sup>44</sup></a><a class="footnote-ref" id="fnref:45" href="#fn:45"><sup>45</sup></a> </p> <h2 id="natural-enemies">Natural enemies</h2> <h3>Predators</h3> <p>There are many predators of the fall webworm, such as <i><a href="/facts/Monomorium_minimum/d55l95ry">Monomorium minimum</a></i>, <i><a href="/facts/Vespula/yV1aldnY">Vespula</a></i> spp., <i><a href="/facts/Coleomegilla_maculata/l74ec11U">Coleomegilla maculata</a></i>, and <i>Rogas hyphantriae</i>.<a class="footnote-ref" id="fnref:46" href="#fn:46"><sup>46</sup></a><a class="footnote-ref" id="fnref:47" href="#fn:47"><sup>47</sup></a> </p> <h3>Parasites</h3> <p>The fall webworm is parasitized by around 50 species, such as the flies <i><a href="/facts/Housefly/GFbeB3In">Musca domestica</a></i> and <i><a href="/facts/Muscina_stabulans/sWW2lEsm">Muscina stabulans</a></i>, and <a href="/facts/Parasitoid_wasp/vLVmVHBs">parasitic wasps</a> such as <i><a href="/facts/Chouioia/UnKXTo2y">Chouioia</a> cunea</i>, <a href="/facts/Meteorus_hyphantriae/FbHUqUeE">Meteorus hyphantriae</a><i>, and </i><a href="/facts/Therion_morio/EJvuIy2V">Therion morio</a><i>.<a class="footnote-ref" id="fnref:48" href="#fn:48"><sup>48</sup></a><a class="footnote-ref" id="fnref:49" href="#fn:49"><sup>49</sup></a><a class="footnote-ref" id="fnref:50" href="#fn:50"><sup>50</sup></a><a class="footnote-ref" id="fnref:51" href="#fn:51"><sup>51</sup></a><a class="footnote-ref" id="fnref:52" href="#fn:52"><sup>52</sup></a> <a class="footnote-ref" id="fnref:53" href="#fn:53"><sup>53</sup></a> There are also several species of <a href="/facts/Microsporidae/fScYY3Il">microsporida</a> that can be harmful, such as </i>Nosema necatrix<i> Kramer (</i>N. necatrix<i>), </i>Pleistophora schubergi hyphantriae<i> Weiser (</i>P.s. hyphantriae<i>), </i><a href="/facts/Nosema_(microsporidian)/nRRwzrUE">Nosema</a><i> sp., and </i><a href="/facts/Nosema_bombycis/nRRwzrUE">Nosema bombycis</a><i> (</i>N. bombycis<i> ).<a class="footnote-ref" id="fnref:54" href="#fn:54"><sup>54</sup></a> </i>Nosema bombycis<i> can affect many body tissues, such as the midgut, fat body, gonads, nervous tissue, and Malpighian tubules, and the length of its polar filament is 100 μm.<a class="footnote-ref" id="fnref:55" href="#fn:55"><sup>55</sup></a> Spores from </i>Nosema<i> of fall webworms are typically egg-shaped and can sometimes be attached to each other, and the length of its polar filament is 57.5 μm ± 8.7 μm.<a class="footnote-ref" id="fnref:56" href="#fn:56"><sup>56</sup></a></i> </p> <h3>Diseases</h3> <p>Several types of pathogens can infect this species, such as <a href="/facts/Granulovirus/E8zeAP9z">granulovirus</a> (which causes granulosis disease) and nucleopolyhedrosis virus.<a class="footnote-ref" id="fnref:57" href="#fn:57"><sup>57</sup></a><a class="footnote-ref" id="fnref:58" href="#fn:58"><sup>58</sup></a><a class="footnote-ref" id="fnref:59" href="#fn:59"><sup>59</sup></a> Granulovirus has a rod-like shape.<a class="footnote-ref" id="fnref:60" href="#fn:60"><sup>60</sup></a> Also, an individual will be more likely to be infected at a younger age.<a class="footnote-ref" id="fnref:61" href="#fn:61"><sup>61</sup></a> </p> <h2 id="interaction-with-humans">Interaction with humans</h2> <p>The fall webworm’s interaction with humans is important because of how adaptable this species is, which is one of the main reasons that it spread so widely across the world.<a class="footnote-ref" id="fnref:62" href="#fn:62"><sup>62</sup></a><a class="footnote-ref" id="fnref:63" href="#fn:63"><sup>63</sup></a> Fall webworms are able to forage on large amounts of leaves inside their web, they are able to create webs on many different types of host plants, they are able to create webs at many different elevations and of many different shapes, and they spread very quickly.<a class="footnote-ref" id="fnref:64" href="#fn:64"><sup>64</sup></a><a class="footnote-ref" id="fnref:65" href="#fn:65"><sup>65</sup></a><a class="footnote-ref" id="fnref:66" href="#fn:66"><sup>66</sup></a><a class="footnote-ref" id="fnref:67" href="#fn:67"><sup>67</sup></a><a class="footnote-ref" id="fnref:68" href="#fn:68"><sup>68</sup></a> In addition, the fall webworm tends to create many webs or defoliate plants in places that humans spend time, such as recreational areas and parks; fall webworms tend to create their webs on branches and therefore defoliate parts of the tree that are very visible and draw attention.<a class="footnote-ref" id="fnref:69" href="#fn:69"><sup>69</sup></a><a class="footnote-ref" id="fnref:70" href="#fn:70"><sup>70</sup></a><a class="footnote-ref" id="fnref:71" href="#fn:71"><sup>71</sup></a><a class="footnote-ref" id="fnref:72" href="#fn:72"><sup>72</sup></a> There have been methods with trying to control this organism, such as physical removal or chemical methods; however, there are some issues with these options because physical removal can be challenging with certain types of trees (based on factors like how tall the tree is, what its function is, where it is located), and the side effects of chemicals (such as important pollinators dying and therefore not being able to perform their functions).<a class="footnote-ref" id="fnref:73" href="#fn:73"><sup>73</sup></a><a class="footnote-ref" id="fnref:74" href="#fn:74"><sup>74</sup></a> New, effective methods are needed to control this organism; Wang <i>et al.</i> (2016) showed the effectiveness of a community-based system to monitor webworms, which involved using mobile devices.<a class="footnote-ref" id="fnref:75" href="#fn:75"><sup>75</sup></a> </p><p>A tree, when covered by the web of the fall webworm is, in the Southern States, sometimes called a "fuzzy tree" because of its fuzzy appearance. </p> <h2 id="further-reading">Further reading</h2> <ul><li>Bat Man, 2003. The fall webworm. <a href="https://web.archive.org/web/20130613053245/http://www.bugwood.org/factsheets/webworm.html">https://web.archive.org/web/20130613053245/http://www.bugwood.org/factsheets/webworm.html</a> . Accessed Aug 21, 2006.</li> <li>Gomi, Takeda (1996). "Changes in life-history traits of Fall Webworm within half a century of introduction into Japan". <i>Functional Ecology</i>. 10 (3): 384–389. <a href="/facts/Doi_(identifier)/muM9Etpq">doi</a>:<a href="https://doi.org/10.2307%2F2390287">10.2307/2390287</a>. <a href="/facts/JSTOR_(identifier)/YTeVmaJ7">JSTOR</a> <a href="https://www.jstor.org/stable/2390287">2390287</a>.</li> <li>Hyche, LL, 1999. Fall webworm: A guide to recognition and habits in Alabama. <a href="http://www.ag.auburn.edu/enpl/bulletins/fallwebworm/fallwebworm.htm">http://www.ag.auburn.edu/enpl/bulletins/fallwebworm/fallwebworm.htm</a> Accessed August 21, 2006.</li> <li>Wagner, DL. 2005. Caterpillars of eastern forests. Princeton Univ. Press.</li> <li>Warren, LO and M Tadic. 1970. The fall webworm, <i>Hyphantria cunea</i> (Drury). Arkansas Agricultural Experiment Station Bulletin 759: 1-106.</li></ul> <h2 id="external-links">External links</h2> Wikimedia Commons has media related to Hyphantria cunea. <a href="/facts/Wikispecies/b2GhA7Le">Wikispecies</a> has information related to <i>Hyphantria cunea</i>. <ul><li><a href="https://www.ukmoths.org.uk/species/hyphantria-cunea/">Fall webworm on UKmoths</a></li> <li><a href="https://web.archive.org/web/20100917225314/http://web.extension.illinois.edu/champaign/homeowners/010818.html">Bagworm, Fall Webworm or Eastern Tent Caterpillar?</a> August 18, 2001. Sandra Mason, University of Illinois Extension. Accessed May 31, 2010.</li> <li><a href="http://www.umassgreeninfo.org/fact_sheets/defoliators/fall_webworm.html">U. Mass site</a> <a href="https://web.archive.org/web/20110928005115/http://www.umassgreeninfo.org/fact_sheets/defoliators/fall_webworm.html">Archived</a> 2011-09-28 at the <a href="/facts/Wayback_Machine/nmQ3a6JC">Wayback Machine</a></li> <li><a href="https://web.archive.org/web/20130613053245/http://www.bugwood.org/factsheets/webworm.html">U. Georgia site</a></li> <li><a href="https://web.archive.org/web/20011006022739/http://ohioline.osu.edu/hyg-fact/2000/2026.html">Ohio State U. site</a></li> <li><a href="https://web.archive.org/web/20050525011347/http://bexar-tx.tamu.edu/HomeHort/F1Column/2004Articles/MAR21.htm">Texas A&M site</a></li> <li><a href="http://entnemdept.ifas.ufl.edu/creatures/trees/moths/fall_webworm.htm">Fall webworm</a> on the <a href="/facts/University_of_Florida/zGg0HXXs">UF</a> / <a href="/facts/Institute_of_Food_and_Agricultural_Sciences/x1JrXu0R">IFAS</a> Featured Creatures website</li></ul> <h2 id="references">References</h2> <ol> <li id="fn:1"><p>Edosa, Tariku T.; Jo, Yong H.; Keshavarz, Maryam; Anh, Young Sang; Noh, Mi Young; Han, Yeon S. (February 2019). "Current status of the management of fall webworm, Hyphantria cunea : Towards the integrated pest management development". Journal of Applied Entomology. 143 (1–2): 1–10. doi:10.1111/jen.12562. S2CID 92540632. <a href="https://doi.org/10.1111%2Fjen.12562" target="_blank">https://doi.org/10.1111%2Fjen.12562</a> <a href="#fnref:1" class="footnote-back-ref">↩</a></p></li> <li id="fn:2"><p>Schowalter, T. D.; Ring, D. R. (2017-01-01). "Biology and Management of the Fall Webworm, Hyphantria cunea (Lepidoptera: Erebidae)". Journal of Integrated Pest Management. 8 (1). doi:10.1093/jipm/pmw019. <a href="https://doi.org/10.1093%2Fjipm%2Fpmw019" target="_blank">https://doi.org/10.1093%2Fjipm%2Fpmw019</a> <a href="#fnref:2" class="footnote-back-ref">↩</a></p></li> <li id="fn:3"><p>Warren, L. O.; Tadić, Milorad (1967). "The Fall Webworm, Hyphantria cunea, Its Distribution and Natural Enemies: A World List (Lepidoptera: Arctiidae)". Journal of the Kansas Entomological Society. 40 (2): 194–202. JSTOR 25083620. <a href="/wiki/JSTOR_(identifier)" target="_blank">/wiki/JSTOR_(identifier)</a> <a href="#fnref:3" class="footnote-back-ref">↩</a></p></li> <li id="fn:4"><p>Lopez-Vaamonde, C.; Agassiz, D.; Augustin, S.; De Prins, J.; De Prins, W.; Gomboc, S.; Ivinskis, P.; Karsholt, O.; Koutroumpas, A.; Koutroumpa, F.; Laštůvka, Z. K.; Marabuto, E.; Olivella, E.; Przybylowicz, L.; Roques, A.; Ryrholm, N.; Sefrova, H.; Sima, P.; Sims, I.; Sinev, S.; Skulev, B.; Tomov, R.; Zilli, A.; Lees, D. (2010). "Lepidoptera. Chapter 11". BioRisk. 4: 603–668. doi:10.3897/biorisk.4.50. <a href="https://doi.org/10.3897%2Fbiorisk.4.50" target="_blank">https://doi.org/10.3897%2Fbiorisk.4.50</a> <a href="#fnref:4" class="footnote-back-ref">↩</a></p></li> <li id="fn:5"><p>Gomi, Takeda (1996). "Changes in life-history traits of Fall Webworm within half a century of introduction into Japan". Functional Ecology. 10 (3): 384–389. doi:10.2307/2390287. JSTOR 2390287. <a href="/wiki/Doi_(identifier)" target="_blank">/wiki/Doi_(identifier)</a> <a href="#fnref:5" class="footnote-back-ref">↩</a></p></li> <li id="fn:6"><p>Edosa, Tariku T.; Jo, Yong H.; Keshavarz, Maryam; Anh, Young Sang; Noh, Mi Young; Han, Yeon S. (February 2019). "Current status of the management of fall webworm, Hyphantria cunea : Towards the integrated pest management development". Journal of Applied Entomology. 143 (1–2): 1–10. doi:10.1111/jen.12562. S2CID 92540632. <a href="https://doi.org/10.1111%2Fjen.12562" target="_blank">https://doi.org/10.1111%2Fjen.12562</a> <a href="#fnref:6" class="footnote-back-ref">↩</a></p></li> <li id="fn:7"><p>Wagner, DL (2005). Caterpillars of Eastern Forests. Princeton, NJ: Princeton Univ. Press. <a href="#fnref:7" class="footnote-back-ref">↩</a></p></li> <li id="fn:8"><p>Richardson, Matthew; Caron, Dewey; Suchanic, David (2006). "Degree-Days for Five Ornamental Pests from an 11-Year Field Study". 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