Related to the Ryūkyū spiny rats (Tokudaia) and the prehistoric Rhagamys – and far more distantly to Mus and Malacomys1 – it includes these species:
Further information: Incertae sedis
Prehistoric species described from fossil remains include:
Steppan et al. (2005) ↩
Alexandra van der Geer; George Lyras; John de Vos; Michael Dermitzakis (2011). Evolution of Island Mammals: Adaptation and Extinction of Placental Mammals on Islands. John Wiley & Sons. ISBN 9781444391282. 9781444391282 ↩