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Clouded leopard
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<h2 id="taxonomy-and-phylogeny">Taxonomy and phylogeny</h2> <p><i>Felis nebulosa</i> was proposed by <a href="/facts/Edward_Griffith_(zoologist)/CY3sHb0H">Edward Griffith</a> in 1821 who first <a href="/facts/Scientific_description/JyFpxSvJ">described</a> a skin of a clouded leopard that was brought alive from <a href="/facts/Guangdong/wvjsjdb1">Guangdong</a> in China to the menagerie at <a href="/facts/Exeter_Exchange/7ccBz2Sf">Exeter Exchange</a> in London.<a class="footnote-ref" id="fnref:1" href="#fn:1"><sup>1</sup></a> <i>Felis macrosceloides</i> proposed by <a href="/facts/Brian_Houghton_Hodgson/HdN90QL1">Brian Houghton Hodgson</a> in 1841 was a clouded leopard <a href="/facts/Zoological_specimen/DPckRhNa">specimen</a> from Nepal.<a class="footnote-ref" id="fnref:2" href="#fn:2"><sup>2</sup></a><a class="footnote-ref" id="fnref:3" href="#fn:3"><sup>3</sup></a> <i><a href="/facts/Formosan_clouded_leopard/WJ4k9moT">Felis brachyura</a></i> proposed by <a href="/facts/Robert_Swinhoe/BeRnULRq">Robert Swinhoe</a> in 1862 was a clouded leopard skin from Taiwan.<a class="footnote-ref" id="fnref:4" href="#fn:4"><sup>4</sup></a> The <a href="/facts/Generic_name_(biology)/m8EFjSms">generic name</a> <i><a href="/facts/Neofelis/AjtB9ABh">Neofelis</a></i> was proposed by <a href="/facts/John_Edward_Gray/6JdrIDH0">John Edward Gray</a> in 1867 who subordinated all three to this <a href="/facts/Genus_(biology)/m8EFjSms">genus</a>.<a class="footnote-ref" id="fnref:5" href="#fn:5"><sup>5</sup></a> At present, <i>N. nebulosa</i> is considered a <a href="/facts/Monotypic_species/QvoM9aGP">monotypic species</a> due to lack of evidence for <a href="/facts/Subspecies/4L9jFacF">subspeciation</a>.<a class="footnote-ref" id="fnref:6" href="#fn:6"><sup>6</sup></a> </p><p><i>Felis diardi</i> proposed by <a href="/facts/Georges_Cuvier/4mmE7NzN">Georges Cuvier</a> in 1823 was based on a clouded leopard skin from Java.<a class="footnote-ref" id="fnref:7" href="#fn:7"><sup>7</sup></a> It was considered a clouded leopard <a href="/facts/Subspecies/4L9jFacF">subspecies</a> by <a href="/facts/Reginald_Innes_Pocock/JzG0MftN">Reginald Innes Pocock</a> in 1917.<a class="footnote-ref" id="fnref:8" href="#fn:8"><sup>8</sup></a> In 2006, it was identified as a distinct <i>Neofelis</i> species, the <a href="/facts/Sunda_clouded_leopard/kA2pPfFE">Sunda clouded leopard</a>.<a class="footnote-ref" id="fnref:9" href="#fn:9"><sup>9</sup></a><a class="footnote-ref" id="fnref:10" href="#fn:10"><sup>10</sup></a> Populations in Taiwan and Hainan Island are considered to belong to the mainland clouded leopard.<a class="footnote-ref" id="fnref:11" href="#fn:11"><sup>11</sup></a> </p> <h3>Phylogeny</h3> <p>Skulls of clouded leopard and <i><a href="/facts/Panthera/qSTyXeae">Panthera</a></i> species were analysed <a href="/facts/Morphology_(biology)/Nl3B0myy">morphologically</a> in the 1960s. Results indicate that the clouded leopard forms an <a href="/facts/Evolution/qE5amP6a">evolutionary</a> link between the <a href="/facts/Pantherinae/r7YKzqKj">Pantherinae</a> and the <a href="/facts/Felinae/pSaoHzNE">Felinae</a>.<a class="footnote-ref" id="fnref:12" href="#fn:12"><sup>12</sup></a> <a href="/facts/Phylogenetic/bw4rsovt">Phylogenetic</a> analysis of the <a href="/facts/Nuclear_DNA/5wzCJQlb">nuclear DNA</a> in tissue samples from all Felidae species revealed that the <a href="/facts/Evolutionary_radiation/DDc0R6GS">evolutionary radiation</a> of the Felidae began in the <a href="/facts/Miocene/cBRRzq4g">Miocene</a> around <a href="https://geoltime.github.io/?Ma=14.45–8.38">14.45 to 8.38</a> million years ago in Asia.<a class="footnote-ref" id="fnref:13" href="#fn:13"><sup>13</sup></a><a class="footnote-ref" id="fnref:14" href="#fn:14"><sup>14</sup></a> Analysis of <a href="/facts/Mitochondrial_DNA/goC0ex0X">mitochondrial DNA</a> of all Felidae species indicates a radiation at <a href="https://geoltime.github.io/?Ma=16.76–6.46">16.76 to 6.46</a> million years ago.<a class="footnote-ref" id="fnref:15" href="#fn:15"><sup>15</sup></a> The clouded leopard is the <a href="/facts/Sister_group/pNNuVex4">sister taxon</a> to all other members of the Pantherinae, diverging <a href="https://geoltime.github.io/?Ma=9.32–4.47">9.32 to 4.47</a> million years ago, based on analysis of their nuclear DNA.<a class="footnote-ref" id="fnref:16" href="#fn:16"><sup>16</sup></a> The clouded leopard from mainland Asia reached <a href="/facts/Borneo/1aIGfyvV">Borneo</a> and <a href="/facts/Sumatra/Fo192kJb">Sumatra</a> via a now submerged <a href="/facts/Land_bridge/WT3Jzmgu">land bridge</a> probably during the <a href="/facts/Pleistocene/0ya9eUp5">Pleistocene</a>, when populations became isolated during periods of global cooling and warming. <a href="/facts/Genetic_analysis/mBppwMUv">Genetic analysis</a> of hair samples of the clouded leopard and its <a href="/facts/Sister_group/pNNuVex4">sister species</a> the Sunda clouded leopard (<i>N. diardi</i>) indicates that they diverged 2.0–0.93 million years ago.<a class="footnote-ref" id="fnref:17" href="#fn:17"><sup>17</sup></a> </p> Phylogenetic relationships of the clouded leopard as derived through analysis of<table><tbody><tr><th>nuclear DNA:<a class="footnote-ref" id="fnref:18" href="#fn:18"><sup>18</sup></a></th></tr><tr><td><table><tbody><tr><td>Felidae</td><td rowspan="2"><table><tbody><tr><td></td><td rowspan="2"><p>Felinae</p></td></tr><tr><td></td></tr><tr><td>Pantherinae</td><td rowspan="2"><table><tbody><tr><td></td><td rowspan="2"><p><i>Panthera</i></p></td></tr><tr><td></td></tr><tr><td></td><td rowspan="2"><p>Clouded leopard</p></td></tr><tr><td></td></tr></tbody></table></td></tr><tr><td></td></tr></tbody></table></td></tr><tr><td></td></tr></tbody></table></td></tr></tbody></table><table><tbody><tr><th>combined mitochondrial and nuclear DNA:<a class="footnote-ref" id="fnref:19" href="#fn:19"><sup>19</sup></a></th></tr><tr><td><table><tbody><tr><td>Felidae</td><td rowspan="2"><table><tbody><tr><td></td><td rowspan="2"><p>Felinae</p></td></tr><tr><td></td></tr><tr><td>Pantherinae</td><td rowspan="2"><table><tbody><tr><td></td><td rowspan="2"><p><i>Panthera</i></p></td></tr><tr><td></td></tr><tr><td><i>Neofelis</i></td><td rowspan="2"><table><tbody><tr><td></td><td rowspan="2"><p>Sunda clouded leopard</p></td></tr><tr><td></td></tr><tr><td></td><td rowspan="2"><p>Mainland clouded leopard</p></td></tr><tr><td></td></tr></tbody></table></td></tr><tr><td></td></tr></tbody></table></td></tr><tr><td></td></tr></tbody></table></td></tr><tr><td></td></tr></tbody></table></td></tr></tbody></table> <h2 id="characteristics">Characteristics</h2> <p>The clouded leopard's fur is of a dark grey or ochreous ground-color, often largely obliterated by black and dark dusky-grey blotched pattern. There are black spots on the head, and the ears are black. Partly fused or broken-up stripes run from the corner of the eyes over the cheek, from the corner of the mouth to the neck, and along the nape to the shoulders. Elongated blotches continue down the spine and form a single median stripe on the loins. Two large blotches of dark dusky-grey hair on the side of the shoulders are each emphasized posteriorly by a dark stripe, which passes on to the foreleg and breaks up into irregular spots. The flanks are marked by dark dusky-grey irregular blotches bordered behind by long, oblique, irregularly curved or looped stripes. These blotches yielding the clouded pattern suggest the English name of the cat. The underparts and legs are spotted, and the tail is marked by large, irregular, paired spots. Its legs are short and stout, and paws broad. Females are slightly smaller than males.<a class="footnote-ref" id="fnref:20" href="#fn:20"><sup>20</sup></a> </p><p>Its <a href="/facts/Hyoid_bone/yAa2Vatb">hyoid bone</a> is ossified, making it possible to purr. Its pupils contract into vertical slits.<a class="footnote-ref" id="fnref:21" href="#fn:21"><sup>21</sup></a> <a href="/facts/Iris_(eye)/e774BXL8">Irises</a> are brownish yellow to grayish green. <a href="/facts/Melanism/PmCo6EEk">Melanistic</a> clouded leopards are uncommon. It has rather short limbs compared to the other big cats. Its hind limbs are longer than its front limbs to allow for increased jumping and leaping capabilities. Its <a href="/facts/Ulnae/6J7D2mhe">ulnae</a> and <a href="/facts/Radius_(bone)/TstRh60d">radii</a> are not fused, which also contributes to a greater range of motion when climbing trees and stalking prey. Clouded leopards weigh between 11.5 and 23 kg (25 and 51 lb). Females vary in head-to-body length from 68.6 to 94 cm (27.0 to 37.0 in), with a tail 61 to 82 cm (24 to 32 in) long. Males are larger at 81 to 108 cm (32 to 43 in) with a tail 74 to 91 cm (29 to 36 in) long.<a class="footnote-ref" id="fnref:22" href="#fn:22"><sup>22</sup></a> Its shoulder height varies from 50 to 55 cm (20 to 22 in).<a class="footnote-ref" id="fnref:23" href="#fn:23"><sup>23</sup></a> </p><p>Its skull is long and low with strong <a href="/facts/Occipital_bone/zsgqpNIa">occipital</a> and <a href="/facts/Sagittal_crest/Mjm0QXDr">sagittal crests</a>. The <a href="/facts/Canine_tooth/p3mRABtg">canine teeth</a> are exceptionally long, the upper being about three times as long as the basal width of the socket. The first premolar is usually absent.<a class="footnote-ref" id="fnref:24" href="#fn:24"><sup>24</sup></a> The upper pair of canines measure 4 cm (1.6 in) or longer.<a class="footnote-ref" id="fnref:25" href="#fn:25"><sup>25</sup></a> It has a <a href="/facts/Bite_force/q1kWzpow">bite force</a> at the <a href="/facts/Canine_tooth/p3mRABtg">canine</a> tip of 544.3 <a href="/facts/Newton_(unit)/Ral9XgNl">Newton</a> and a <a href="/facts/Bite_force_quotient/q1kWzpow">bite force quotient</a> at the canine tip of 122.4.<a class="footnote-ref" id="fnref:26" href="#fn:26"><sup>26</sup></a> The clouded leopard is often referred to as a "modern-day <a href="/facts/Saber-toothed_cat/29SyW7vn">sabre-tooth</a>" because it has the largest canines in proportion to its body size.<a class="footnote-ref" id="fnref:27" href="#fn:27"><sup>27</sup></a> </p> <h2 id="distribution-and-habitat">Distribution and habitat</h2> <p>The clouded leopard occurs from the <a href="/facts/Himalayas/eRSf0yyE">Himalayan</a> foothills in <a href="/facts/Nepal/yWCzTBR2">Nepal</a>, <a href="/facts/Bhutan/fSRsLss9">Bhutan</a> and <a href="/facts/India/QW4TV4sy">India</a> to <a href="/facts/Myanmar/zcqQH2oE">Myanmar</a>, southeastern <a href="/facts/Bangladesh/NXk4f78Y">Bangladesh</a>, <a href="/facts/Thailand/GCUnN4JK">Thailand</a>, <a href="/facts/Peninsular_Malaysia/88WtYx31">Peninsular Malaysia</a> and to south of the <a href="/facts/Yangtze_River/f9A2h4rQ">Yangtze River</a> in China. It is locally extinct in Singapore and Taiwan.<a class="footnote-ref" id="fnref:28" href="#fn:28"><sup>28</sup></a> </p><p>Clouded leopards were found in Nepal in 1987 and 1988, having previously been presumed to be extinct in the country.<a class="footnote-ref" id="fnref:29" href="#fn:29"><sup>29</sup></a> Since then, the clouded leopard has been recorded in <a href="/facts/Shivapuri_Nagarjun_National_Park/24gFaG4Y">Shivapuri Nagarjun National Park</a> and in <a href="/facts/Annapurna_Conservation_Area/enpqbFHN">Annapurna Conservation Area</a>.<a class="footnote-ref" id="fnref:30" href="#fn:30"><sup>30</sup></a><a class="footnote-ref" id="fnref:31" href="#fn:31"><sup>31</sup></a> Between 2014 and 2015, it was also recorded in <a href="/facts/Langtang_National_Park/fkjDVKz8">Langtang National Park</a> at an elevation range of 1,823–3,498 m (5,981–11,476 ft).<a class="footnote-ref" id="fnref:32" href="#fn:32"><sup>32</sup></a> </p><p>In India, it occurs in the states of <a href="/facts/Sikkim/x89Pn6xf">Sikkim</a>, northern <a href="/facts/West_Bengal/0B5bxvbW">West Bengal</a>, <a href="/facts/Tripura/zbaay5BU">Tripura</a>, <a href="/facts/Mizoram/RQXMgMFl">Mizoram</a>, <a href="/facts/Manipur/afIbV0R2">Manipur</a>, <a href="/facts/Assam/64Hs7TwB">Assam</a>, <a href="/facts/Nagaland/DIhsVRqL">Nagaland</a> and <a href="/facts/Arunachal_Pradesh/4rrSKgGE">Arunachal Pradesh</a>, as well as in the <a href="/facts/Meghalaya_subtropical_forests/vzmLgncX">Meghalaya subtropical forests</a>.<a class="footnote-ref" id="fnref:33" href="#fn:33"><sup>33</sup></a><a class="footnote-ref" id="fnref:34" href="#fn:34"><sup>34</sup></a><a class="footnote-ref" id="fnref:35" href="#fn:35"><sup>35</sup></a><a class="footnote-ref" id="fnref:36" href="#fn:36"><sup>36</sup></a> In <a href="/facts/Pakke_Tiger_Reserve/nbIjnz7v">Pakke Tiger Reserve</a>, a clouded leopard was photographed in <a href="/facts/Semi-evergreen/AJ5HXnYR">semi-evergreen</a> forest at an elevation of 144 m (472 ft).<a class="footnote-ref" id="fnref:37" href="#fn:37"><sup>37</sup></a> In Sikkim, clouded leopards were photographed by <a href="/facts/Camera_trap/uU0TFx4k">camera traps</a> at elevations of 2,500–3,720 m (8,200–12,200 ft) between April 2008 and May 2010 in the <a href="/facts/Khangchendzonga_National_Park/Qg1N2d99">Khangchendzonga Biosphere Reserve</a>.<a class="footnote-ref" id="fnref:38" href="#fn:38"><sup>38</sup></a> In <a href="/facts/Manas_National_Park/gMffHnmj">Manas National Park</a>, 16 individuals were recorded during a survey in November 2010 to February 2011.<a class="footnote-ref" id="fnref:39" href="#fn:39"><sup>39</sup></a> Between January 2013 and March 2018, clouded leopards were also recorded in <a href="/facts/Dampa_Tiger_Reserve/vWQjkKtL">Dampa Tiger Reserve</a>, <a href="/facts/Eaglenest_Wildlife_Sanctuary/pTUB6oo6">Eaglenest Wildlife Sanctuary</a> and Singchung-Bugun Village Community Reserve, in Meghalaya's Nongkhyllem National Park and <a href="/facts/Balpakram/rgAsdXV7">Balpakram</a>-<a href="/facts/Baghmara%2c_India/715J0Rwy">Baghmara</a> landscape.<a class="footnote-ref" id="fnref:40" href="#fn:40"><sup>40</sup></a> </p><p>In Bhutan, it was recorded in <a href="/facts/Royal_Manas_National_Park/blGcDNke">Royal Manas National Park</a>, <a href="/facts/Jigme_Singye_Wangchuck_National_Park/pyrrW71g">Jigme Singye Wangchuck National Park</a>, <a href="/facts/Phibsoo_Wildlife_Sanctuary/xHrEw5hG">Phibsoo Wildlife Sanctuary</a>, <a href="/facts/Jigme_Dorji_National_Park/ARGwDsaa">Jigme Dorji National Park</a>, <a href="/facts/Phrumsengla_National_Park/GzsbnQsl">Phrumsengla National Park</a>, <a href="/facts/Bumdeling_Wildlife_Sanctuary/NgReNYC3">Bumdeling Wildlife Sanctuary</a> and several non-protected areas.<a class="footnote-ref" id="fnref:41" href="#fn:41"><sup>41</sup></a><a class="footnote-ref" id="fnref:42" href="#fn:42"><sup>42</sup></a> In Bangladesh, it was recorded in <a href="/facts/Sangu_Matamuhari/x8WmX3st">Sangu Matamuhari</a> in the <a href="/facts/Chittagong_Hill_Tracts/gDCNvQsI">Chittagong Hill Tracts</a> in 2016.<a class="footnote-ref" id="fnref:43" href="#fn:43"><sup>43</sup></a> In Myanmar, it was recorded by camera traps for the first time in the hill forests of <a href="/facts/Karen_State/IVg0HySg">Karen State</a> in 2015.<a class="footnote-ref" id="fnref:44" href="#fn:44"><sup>44</sup></a> </p><p>In Thailand, it inhabits relatively open, <a href="/facts/Dry_tropical_forest/o88ufPT5">dry tropical forest</a> in <a href="/facts/Huai_Kha_Khaeng_Wildlife_Sanctuary/KRHxAFX2">Huai Kha Khaeng Wildlife Sanctuary</a> and closed-forest <a href="/facts/Habitat/zo1DN2gA">habitats</a> in <a href="/facts/Khao_Yai_National_Park/Emv0K3o7">Khao Yai National Park</a>.<a class="footnote-ref" id="fnref:45" href="#fn:45"><sup>45</sup></a><a class="footnote-ref" id="fnref:46" href="#fn:46"><sup>46</sup></a><a class="footnote-ref" id="fnref:47" href="#fn:47"><sup>47</sup></a> In Laos, it was recorded in <a href="/facts/Nam_Et-Phou_Louey/JI9gWUI7">Nam Et-Phou Louey</a> National Protected Area in dry <a href="/facts/Evergreen_forest/MFmRRUcR">evergreen</a> and semi-evergreen forests.<a class="footnote-ref" id="fnref:48" href="#fn:48"><sup>48</sup></a> In Cambodia, it was recorded in deciduous <a href="/facts/Dipterocarp_forest/GDUkWoUr">dipterocarp forest</a> in <a href="/facts/Phnom_Prich_Wildlife_Sanctuary/5fsVI1E7">Phnom Prich Wildlife Sanctuary</a> between 2008 and 2009,<a class="footnote-ref" id="fnref:49" href="#fn:49"><sup>49</sup></a> and in <a href="/facts/Central_Cardamom_Mountains_National_Park/UKHBeb9W">Central Cardamom Mountains National Park</a>, <a href="/facts/Southern_Cardamom_National_Park/kNeyLztl">Southern Cardamom National Park</a>, <a href="/facts/Botum_Sakor_National_Park/H4e1CdPY">Botum Sakor National Park</a> and <a href="/facts/Phnom_Samkos_Wildlife_Sanctuary/gtIeyvTe">Phnom Samkos Wildlife Sanctuary</a> between 2012 and 2016.<a class="footnote-ref" id="fnref:50" href="#fn:50"><sup>50</sup></a> In Peninsular Malaysia, it was recorded in <a href="/facts/Taman_Negara_National_Park/aBA8bUL4">Taman Negara National Park</a>, <a href="/facts/Ulu_Muda_Forest/wmeycnLU">Ulu Muda Forest</a>, <a href="/facts/Pasoh_Forest_Reserve/yu1x4B8e">Pasoh Forest Reserve</a>, <a href="/facts/Belum-Temengor/8HqgbMe5">Belum-Temengor</a>, Temengor Forest Reserve and in a few linkages between 2009 and 2015.<a class="footnote-ref" id="fnref:51" href="#fn:51"><sup>51</sup></a> </p><p>The last confirmed record of a Formosan clouded leopard dates to 1989, when the skin of a young individual was found in the <a href="/facts/Taroko_National_Park/nS4r9omq">Taroko National Park</a>.<a class="footnote-ref" id="fnref:52" href="#fn:52"><sup>52</sup></a> It was not recorded during an extensive camera trapping survey conducted from 1997 to 2012 in more than 1,450 sites inside and outside Taiwanese protected areas.<a class="footnote-ref" id="fnref:53" href="#fn:53"><sup>53</sup></a> </p> <h2 id="behaviour-and-ecology">Behaviour and ecology</h2> <p>The clouded leopard is a solitary cat. Early accounts depict it as a rare, secretive, <a href="/facts/Arboreal/6C5AeQ2y">arboreal</a>, and <a href="/facts/Nocturnal/PqAkbpKf">nocturnal</a> inhabitant of dense <a href="/facts/Primary_forest/kffLtWkA">primary forest</a>.<a class="footnote-ref" id="fnref:54" href="#fn:54"><sup>54</sup></a> </p><p>It is one of the most talented climbers among the cats. Captive clouded leopards have been observed to climb down vertical tree trunks head first, and hang on to branches with their hind paws bent around branchings of tree limbs.<a class="footnote-ref" id="fnref:55" href="#fn:55"><sup>55</sup></a> They are capable of <a href="/facts/Supination/KmBVfcPk">supination</a> and can hang down from branches only by bending their hind paws and their tail around them. They can jump up to 1.2 m (3 ft 11 in) high.<a class="footnote-ref" id="fnref:56" href="#fn:56"><sup>56</sup></a> They use trees as daytime rest sites, but also spend time on the ground when hunting at night. Captive clouded leopards have been observed to <a href="/facts/Scent_mark/T15S7XeD">scent mark</a> by <a href="/facts/Urine_spraying/H4qYz02K">spraying urine</a> and <a href="/facts/Scent_rubbing/Nudp04gu">rubbing their heads</a> on prominent objects.<a class="footnote-ref" id="fnref:57" href="#fn:57"><sup>57</sup></a> Their vocalisations include a short high-pitched meow call, a loud crying call, both emitted when a cat is trying to locate another one over a long or short distance; they <a href="/facts/Prusten/0nRVNulU">prusten</a> and raise their muzzle when meeting each other in a friendly manner; when aggressive, they growl with a low-pitched sound and hiss with exposed teeth and wrinkled nose.<a class="footnote-ref" id="fnref:58" href="#fn:58"><sup>58</sup></a> </p><p>Radio-collared clouded leopards were foremost active by night but also showed <a href="/facts/Crepuscular/nHEUNhSo">crepuscular</a> activity peaks.<a class="footnote-ref" id="fnref:59" href="#fn:59"><sup>59</sup></a> Clouded leopards recorded in northeast India were most active in the late evening after sunset.<a class="footnote-ref" id="fnref:60" href="#fn:60"><sup>60</sup></a> </p><p>Home ranges have only been estimated in Thailand: </p> <ul><li>Four individuals were <a href="/facts/Radio_telemetry/Mz9czXTl">radio-collared</a> in Phu Khieo Wildlife Sanctuary from April 2000 to February 2003. Home ranges of two females were 25.7 km2 (9.9 sq mi) and 22.9 km2 (8.8 sq mi), and of two males 29.7 km2 (11.5 sq mi) and 49.1 km2 (19.0 sq mi).<a class="footnote-ref" id="fnref:61" href="#fn:61"><sup>61</sup></a></li> <li>Two individuals were radio-collared during a study from 1997 to 1999 in the <a href="/facts/Khao_Yai_National_Park/Emv0K3o7">Khao Yai National Park</a>. The home range of one female was 39.4 km2 (15.2 sq mi), of the one male 42 km2 (16 sq mi). Both individuals had a core area of 2.9 km2 (1.1 sq mi).<a class="footnote-ref" id="fnref:62" href="#fn:62"><sup>62</sup></a></li></ul> <p>In 2016, clouded leopards were detected in the forest complex of <a href="/facts/Khlong_Saeng_Wildlife_Sanctuary/lVdl5lMH">Khlong Saeng Wildlife Sanctuary</a> and <a href="/facts/Khao_Sok_National_Park/lbNK6uqf">Khao Sok National Park</a> during camera trapping surveys; 15 individuals were identified in a core zone of 200 km2 (77 sq mi) with <a href="/facts/Population_density/70zGTCFc">population density</a> estimated at 5.06 individuals per 100 km2 (39 sq mi); but only 12 individuals were identified in an edge zone of 297 km2 (115 sq mi), which is more disturbed by humans, with density estimated at 3.13 individuals per 100 km2 (39 sq mi).<a class="footnote-ref" id="fnref:63" href="#fn:63"><sup>63</sup></a> </p> <h3>Hunting and diet</h3> <p>When hunting, the clouded leopard stalks its prey or waits for the prey to approach. After making and feeding on a kill, it usually retreats into trees to digest and rest. Its prey includes both arboreal and <a href="/facts/Terrestrial_animal/AhP2mont">terrestrial</a> <a href="/facts/Vertebrate/tT54FSTT">vertebrates</a>.<a class="footnote-ref" id="fnref:64" href="#fn:64"><sup>64</sup></a> Pocock presumed that it is adapted for preying upon <a href="/facts/Herbivore/me4YFdQA">herbivorous</a> <a href="/facts/Mammals/phSwTzBo">mammals</a> of considerable bulk because of its powerful build, long canines and the deep penetration of its bites.<a class="footnote-ref" id="fnref:65" href="#fn:65"><sup>65</sup></a> In Thailand, clouded leopards have been observed preying on <a href="/facts/Southern_pig-tailed_macaque/1jTNwFht">southern pig-tailed macaque</a> (<i>Macaca nemestrina</i>),<a class="footnote-ref" id="fnref:66" href="#fn:66"><sup>66</sup></a> <a href="/facts/Indian_hog_deer/bfThRf6h">Indian hog deer</a> (<i>Axis porcinus</i>), <a href="/facts/Bengal_slow_loris/2VWWnvGg">Bengal slow loris</a> (<i>Nycticebus bengalensis</i>), <a href="/facts/Asiatic_brush-tailed_porcupine/P9T4KXX7">Asiatic brush-tailed porcupine</a> (<i>Atherurus macrourus</i>), <a href="/facts/Malayan_pangolin/myvg2wmB">Malayan pangolin</a> (<i>Manis javanica</i>) and <a href="/facts/Berdmore%2527s_ground_squirrel/pJNtRgpE">Berdmore's ground squirrel</a> (<i>Menetes berdmorei</i>).<a class="footnote-ref" id="fnref:67" href="#fn:67"><sup>67</sup></a> Known prey species in China include <a href="/facts/Barking_deer/0kYy2IRv">barking deer</a> (<i>Muntiacus</i> sp.) and <a href="/facts/Pheasants/TS49HPgs">pheasants</a>.<a class="footnote-ref" id="fnref:68" href="#fn:68"><sup>68</sup></a> In northern Peninsular Malaysia, a male clouded leopard was photographed while carrying a <a href="/facts/Binturong/C87tS5sd">binturong</a> (<i>Arctictis binturong</i>) in its jaws.<a class="footnote-ref" id="fnref:69" href="#fn:69"><sup>69</sup></a> </p> <h3>Reproduction and life cycle</h3> <p>Both males and females average 26 months at first reproduction. The female is in <a href="/facts/Estrus/qo2BuxlQ">estrus</a> for about six days, with her <a href="/facts/Estrous_cycle/qo2BuxlQ">estrous cycle</a> lasting about 30 days.<a class="footnote-ref" id="fnref:70" href="#fn:70"><sup>70</sup></a> In the wild, <a href="/facts/Mating/o2voslGF">mating</a> usually occurs between December and March. The pair mates multiple times over the course of several days. The male grasps the female by the neck who responds with vocalization. Occasionally, he also bites her during courtship and is very aggressive during sexual encounters. Females can bear one <a href="/facts/Litter_(zoology)/ttjkfqWS">litter</a> each year. The male is not involved in raising the cubs.<a class="footnote-ref" id="fnref:71" href="#fn:71"><sup>71</sup></a> </p><p>The female gives birth to a litter of one to five, mostly three cubs, after a <a href="/facts/Gestation/QKgkykw5">gestation</a> period of 93 ± 6 days.<a class="footnote-ref" id="fnref:72" href="#fn:72"><sup>72</sup></a> Cubs are born with closed eyes and weigh from 140 to 280 g (4.9 to 9.9 oz). Their spots are solid dark, rather than dark rings. Their eyes open after about 10 days. They are active within five weeks and fully weaned at around three months of age. They attain the adult coat pattern at around six months and become independent after around 10 months.<a class="footnote-ref" id="fnref:73" href="#fn:73"><sup>73</sup></a> </p><p>Captive clouded leopards have an average lifespan of 11 years.<a class="footnote-ref" id="fnref:74" href="#fn:74"><sup>74</sup></a> One individual has lived to be almost 17 years old.<a class="footnote-ref" id="fnref:75" href="#fn:75"><sup>75</sup></a> </p><p>The <a href="/facts/Generation_time/J7m8p79J">generation length</a> of the clouded leopard is about seven years.<a class="footnote-ref" id="fnref:76" href="#fn:76"><sup>76</sup></a> </p> <h2 id="threats">Threats</h2> <p>Clouded leopard require larger areas of intact forest than are present in many parts of their range.<a class="footnote-ref" id="fnref:77" href="#fn:77"><sup>77</sup></a> They are threatened by habitat loss following large–scale <a href="/facts/Deforestation/L1TNhN9p">deforestation</a> and commercial poaching for the <a href="/facts/Wildlife_trade/WNLPEduI">wildlife trade</a>.<a class="footnote-ref" id="fnref:78" href="#fn:78"><sup>78</sup></a> In Myanmar, 301 body parts of at least 279 clouded leopards, mostly skins and skeletons, were observed in four markets surveyed between 1991 and 2006, despite the protected status of clouded leopards in Myanmar. Some markets are located near Myanmar's borders with China and Thailand and are used to facilitate cross-border smuggling.<a class="footnote-ref" id="fnref:79" href="#fn:79"><sup>79</sup></a> </p><p>In Nepal, 27 cases of clouded leopard body parts were discovered between November 1988 and March 2020 in nine districts of the country, comprising at least 51 individual clouded leopards. In 17 of these cases, the poachers and traders were arrested.<a class="footnote-ref" id="fnref:80" href="#fn:80"><sup>80</sup></a> </p> <h2 id="conservation">Conservation</h2> <p>The clouded leopard is listed in <a href="/facts/CITES_Appendix_I/hCahxMSr">CITES Appendix I</a>. Hunting is banned in Bangladesh, China, India, <a href="/facts/Malaysia/8zbUMzCP">Malaysia</a>, Myanmar, Nepal, Taiwan, Thailand and Vietnam.<a class="footnote-ref" id="fnref:81" href="#fn:81"><sup>81</sup></a> These bans, however, are poorly enforced in India, Malaysia and Thailand.<a class="footnote-ref" id="fnref:82" href="#fn:82"><sup>82</sup></a> </p><p>In the <a href="/facts/United_States/P0sCdLe5">United States</a>, the clouded leopard is listed as <a href="/facts/Endangered_species/L7cIweSW">endangered</a> under the <a href="/facts/Endangered_Species_Act/voufDnBj">Endangered Species Act</a>, prohibiting trade in live animals or body parts.<a class="footnote-ref" id="fnref:83" href="#fn:83"><sup>83</sup></a> </p><p>International Clouded Leopard Day is celebrated each year on 4 August since 2018 in zoos and conservation organizations all over the world.<a class="footnote-ref" id="fnref:84" href="#fn:84"><sup>84</sup></a> </p> <h3>In captivity</h3> <p>Clouded leopards have been kept in zoos since the early 20th century. The international <a href="/facts/Studbook/TV2ApgCg">studbook</a> was initiated in the 1970s. Coordinated breeding programs were started in the 1980s and encompass the <a href="/facts/European_Endangered_Species_Programme/QyMKwd3o">European Endangered Species Programme</a>, the <a href="/facts/Species_Survival_Plan/KJVKG8bX">Species Survival Plan</a>, and the Indian Conservation Breeding Programme. As of 2014, 64 institutions keep clouded leopards.<a class="footnote-ref" id="fnref:85" href="#fn:85"><sup>85</sup></a> </p><p>Early captive-breeding programs involving clouded leopards were not successful, largely due to ignorance of their courtship behaviour. Males have the reputation of being aggressive towards females. For breeding success, it has been deemed extremely important that male and female clouded leopards are compatible.<a class="footnote-ref" id="fnref:86" href="#fn:86"><sup>86</sup></a> Introducing pairs at a young age gives them opportunities to bond and breed successfully. Facilities breeding clouded leopards need to provide the female with a secluded, off-exhibit area.<a class="footnote-ref" id="fnref:87" href="#fn:87"><sup>87</sup></a> There has been some recent captive breeding success using <a href="/facts/Artificial_insemination/ccbKoyu4">artificial insemination</a> with cubs successfully born in 1992, 2015 and 2017.<a class="footnote-ref" id="fnref:88" href="#fn:88"><sup>88</sup></a> </p><p>A study on <a href="/facts/Morbidity/55eeQlGL">morbidity</a> and <a href="/facts/Mortality_rate/s4gmxyFV">mortality rate</a> of 271 captive clouded leopards across 44 zoos in Europe, Asia and Australia showed that 17% of them died because of <a href="/facts/Respiratory_disease/CDRTg5Ev">respiratory disease</a>, 12% due to maternal neglect and starvation, 10% from generalized infectious disease, 10% from <a href="/facts/Digestive_disease/S4UzQRFA">digestive diseases</a>, and 10% from trauma.<a class="footnote-ref" id="fnref:89" href="#fn:89"><sup>89</sup></a> </p><p>In March 2011, two breeding females at the <a href="/facts/Nashville_Zoo_at_Grassmere/jL4GuwBD">Nashville Zoo at Grassmere</a> gave birth to three cubs, which were raised by zookeepers. Each cub weighed 0.23 kg (0.5 lb).<a class="footnote-ref" id="fnref:90" href="#fn:90"><sup>90</sup></a> In June 2011, two cubs were born at the <a href="/facts/Point_Defiance_Zoo_%2526_Aquarium/1AUkM8m6">Point Defiance Zoo & Aquarium</a>. The breeding pair was brought from the <a href="/facts/Khao_Kheow_Open_Zoo/JRbwsSjb">Khao Kheow Open Zoo</a> in Thailand in an ongoing education and research exchange program.<a class="footnote-ref" id="fnref:91" href="#fn:91"><sup>91</sup></a> Four cubs were born at Nashville Zoo in 2012.<a class="footnote-ref" id="fnref:92" href="#fn:92"><sup>92</sup></a> In May 2015, four cubs were born in Point Defiance Zoo & Aquarium.<a class="footnote-ref" id="fnref:93" href="#fn:93"><sup>93</sup></a> </p> <h2 id="in-culture">In culture</h2> <p>The clouded leopard is the state animal of the Indian state of <a href="/facts/Meghalaya/lkm1Upzw">Meghalaya</a>.<a class="footnote-ref" id="fnref:94" href="#fn:94"><sup>94</sup></a> In the 1970s, the print of <a href="/facts/Rama_Samaraweera/jolK9ByQ">Rama Samaraweera</a>'s painting <i>Clouded leopard</i> was a best-seller in the US.<a class="footnote-ref" id="fnref:95" href="#fn:95"><sup>95</sup></a> </p> <h2 id="see-also">See also</h2> <ul><li><a href="/facts/List_of_largest_cats/4GeIBblT">List of largest cats</a></li></ul> <h2 id="external-links">External links</h2> <a href="/facts/Wikimedia_Commons/lzkBFxTg">Wikimedia Commons</a> has media related to: Neofelis nebulosa (category) <a href="/facts/Wikispecies/b2GhA7Le">Wikispecies</a> has information related to <i>Neofelis nebulosa</i>. <ul><li><a href="http://www.catsg.org/index.php?id=116">"Mainland clouded leopard <i>Neofelis nebulosa</i>"</a>. <i>IUCN Species Survival Commission Cat Specialist Group</i>.</li> <li><a href="https://web.archive.org/web/20120224232054/http://animals.nationalgeographic.com/animals/mammals/clouded-leopard.html">"Clouded Leopard <i>Neofelis nebulosa</i>"</a>. <i><a href="/facts/National_Geographic_(magazine)/vnuhsmYg">National Geographic</a></i>. Archived from <a href="http://animals.nationalgeographic.com/animals/mammals/clouded-leopard.html">the original</a> on 24 February 2012.</li> <li><a href="https://focustaiwan.tw/society/201304300027">"Taiwan's clouded leopard extinct: zoologists"</a>. <i>Focus Taiwan</i>. 2013.</li> <li><a href="https://www.aboutanimals.com/mammal/clouded-leopard/">"Clouded Leopard: Facing Dark Clouds of Extinction"</a>. <i>About Animals</i>. 20 June 2016.</li></ul> <h2 id="references">References</h2> <ol> <li id="fn:1"><p>Griffith, E. (1821). "The Chinese, or tortoiseshell Tiger. Felis nebulosa". General and particular descriptions of the vertebrated animals arranged comfortably to the modern discoveries and improvements in zoology. London: Baldwin, Cradock & Joy. p. 37. Archived from the original on 10 February 2023. 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