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Anoual Formation
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<h2 id="paleobiota">Paleobiota</h2> <h3>Brachiopoda</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i>Burmirhynchia</i><a class="footnote-ref" id="fnref:3" href="#fn:3"><sup>3</sup></a></td><td><ul><li><i>B. gr. termierae-athiaensis</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated shells</td><td>A Branchiopod of the family Tetrarhynchiidae</td><td></td></tr><tr><td><i><a href="/facts/Kallirhynchia/1CeJPCVU">Kallirhynchia</a></i><a class="footnote-ref" id="fnref:4" href="#fn:4"><sup>4</sup></a></td><td><ul><li><i>K. oranensis</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated shells</td><td>A Branchiopod of the family Tetrarhynchiidae</td><td></td></tr></tbody></table> <h3>Crustacea</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i>Asciocythere</i><a class="footnote-ref" id="fnref:5" href="#fn:5"><sup>5</sup></a></td><td><ul><li><i>A. sp.</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated Valves</td><td>A Marine <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family Schulerideidae.</td><td></td></tr><tr><td><i>Bythocypris</i><a class="footnote-ref" id="fnref:6" href="#fn:6"><sup>6</sup></a></td><td><ul><li><i>B. sp.</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated Valves</td><td>A Marine <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family <a href="/facts/Bythocyprididae/cPh8pkgT">Bythocyprididae</a></td><td></td></tr><tr><td><i>Asciocythere</i><a class="footnote-ref" id="fnref:7" href="#fn:7"><sup>7</sup></a></td><td><ul><li><i>A. sp.</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated Valves</td><td>A Marine <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family Schulerideidae.</td><td></td></tr><tr><td><i>Cypridea</i><a class="footnote-ref" id="fnref:8" href="#fn:8"><sup>8</sup></a></td><td><ul><li><i>C. cf. postelongata</i></li></ul></td><td>Anoual</td><td>Lower Member</td><td>Isolated Valves</td><td>A Freshwater <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family <a href="/facts/Cyprididae/rbIrB0Mk">Cyprididae</a></td><td></td></tr><tr><td><i>Cypris</i><a class="footnote-ref" id="fnref:9" href="#fn:9"><sup>9</sup></a></td><td><ul><li><i>C. spp.</i></li></ul></td><td>Anoual</td><td>Lower Member</td><td>Isolated Valves</td><td>A Freshwater <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family <a href="/facts/Cyprididae/rbIrB0Mk">Cyprididae</a></td><td></td></tr><tr><td><i><a href="/facts/Darwinula/WjmxYTfw">Darwinula</a></i><a class="footnote-ref" id="fnref:10" href="#fn:10"><sup>10</sup></a></td><td><ul><li><i>D. magna</i></li><li><i>D. cf. leguminella</i></li></ul></td><td>Anoual</td><td>Lower Member</td><td>Isolated Valves</td><td>A Freshwater <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family <a href="/facts/Darwinulidae/m9URMJJC">Darwinulidae</a></td><td></td></tr><tr><td><i>Fabanella</i><a class="footnote-ref" id="fnref:11" href="#fn:11"><sup>11</sup></a></td><td><ul><li><i>F. bathonica</i></li><li><i>F. sp.</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated Valves</td><td>A Freshwater <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the order <a href="/facts/Podocopida/ntfFplk1">Podocopida</a></td><td></td></tr><tr><td><i>Fastigocythere</i><a class="footnote-ref" id="fnref:12" href="#fn:12"><sup>12</sup></a></td><td><ul><li><i>F. sp.</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated Valves</td><td>A Marine <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the order <a href="/facts/Podocopida/ntfFplk1">Podocopida</a></td><td></td></tr><tr><td><i>Klieana</i><a class="footnote-ref" id="fnref:13" href="#fn:13"><sup>13</sup></a></td><td><ul><li><i>K. levis</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated Valves</td><td>A Freshwater <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the order <a href="/facts/Cytherideidae/lfphpIPJ">Cytherideidae</a></td><td></td></tr><tr><td><i>Kinkelinella</i><a class="footnote-ref" id="fnref:14" href="#fn:14"><sup>14</sup></a></td><td><ul><li><i>K. sp.</i></li></ul></td><td>Anoual</td><td>Lower Member</td><td>Isolated Valves</td><td>A Freshwater/Brackish <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family <a href="/facts/Protocytheridae/pg5UDI4u">Protocytheridae</a></td><td></td></tr><tr><td><i>Mantelliana</i><a class="footnote-ref" id="fnref:15" href="#fn:15"><sup>15</sup></a></td><td><ul><li><i>M. sp.</i></li></ul></td><td>Anoual</td><td>Lower Member</td><td>Isolated Valves</td><td>A Freshwater <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family <a href="/facts/Cyprididae/rbIrB0Mk">Cyprididae</a></td><td></td></tr><tr><td><i><a href="/facts/Metacypris/V7WP1fNb">Metacypris</a></i><a class="footnote-ref" id="fnref:16" href="#fn:16"><sup>16</sup></a></td><td><ul><li><i>M. sp.</i></li></ul></td><td>Anoual</td><td>Lower Member</td><td>Isolated Valves</td><td>A Freshwater/Brackish <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family <a href="/facts/Limnocytheridae/MLYDcRqp">Limnocytheridae</a></td><td></td></tr><tr><td><i>Paracypris</i><a class="footnote-ref" id="fnref:17" href="#fn:17"><sup>17</sup></a></td><td><ul><li><i>P. sp.</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated Valves</td><td>A Marine <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family <a href="/facts/Candonidae/b8uC6W76">Candonidae</a>. Dominant marine taxon</td><td></td></tr><tr><td><i>Pneumatocythere</i><a class="footnote-ref" id="fnref:18" href="#fn:18"><sup>18</sup></a></td><td><ul><li><i>P.? juglandiformis</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated Valves</td><td>A Marine <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the order <a href="/facts/Podocopida/ntfFplk1">Podocopida</a></td><td></td></tr><tr><td><i>Praeschuleridea</i><a class="footnote-ref" id="fnref:19" href="#fn:19"><sup>19</sup></a></td><td><ul><li><i>P. sp.</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated Valves</td><td>A Marine <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family <a href="/facts/Candonidae/b8uC6W76">Candonidae</a>.</td><td></td></tr><tr><td><i>Procytheridea</i><a class="footnote-ref" id="fnref:20" href="#fn:20"><sup>20</sup></a></td><td><ul><li><i>P. sp.</i></li></ul></td><td>Anoual</td><td>Upper Member</td><td>Isolated Valves</td><td>A Marine <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the order <a href="/facts/Podocopida/ntfFplk1">Podocopida</a></td><td></td></tr><tr><td><i>Scabriculocypris</i><a class="footnote-ref" id="fnref:21" href="#fn:21"><sup>21</sup></a></td><td><ul><li><i>S. sp.</i></li></ul></td><td>Anoual</td><td>Lower Member</td><td>Isolated Valves</td><td>A Freshwater <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family <a href="/facts/Cyprididae/rbIrB0Mk">Cyprididae</a></td><td></td></tr><tr><td><i>Theriosynoecum</i><a class="footnote-ref" id="fnref:22" href="#fn:22"><sup>22</sup></a></td><td><ul><li><i>T. sp.</i></li></ul></td><td>Anoual</td><td>Lower Member</td><td>Isolated Valves</td><td>A Freshwater/Brackish <a href="/facts/Ostracod/q7ejFS5I">Ostracod</a> of the family <a href="/facts/Limnocytheridae/MLYDcRqp">Limnocytheridae</a></td><td></td></tr></tbody></table> <h3>Mollusca</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><a href="/facts/Lymnaeidae/tQCq2Ra6">Lymnaeidae</a><a class="footnote-ref" id="fnref:23" href="#fn:23"><sup>23</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Isolated shells</td><td>Freshwater snails</td><td></td></tr><tr><td><a href="/facts/Nerineidae/krNCTUVw">Nerineidae</a><a class="footnote-ref" id="fnref:24" href="#fn:24"><sup>24</sup></a></td><td>Indeterminate</td><td>Anoual</td><td>Upper Member</td><td>Isolated shells</td><td>Marine snails</td><td></td></tr><tr><td><a href="/facts/Planorbidae/UIISsvMB">Planorbidae</a><a class="footnote-ref" id="fnref:25" href="#fn:25"><sup>25</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Isolated shells</td><td>Freshwater snails</td><td></td></tr><tr><td><i>Provalvata</i><a class="footnote-ref" id="fnref:26" href="#fn:26"><sup>26</sup></a></td><td><i>P. sp.</i></td><td>Anoual</td><td>Lower Member</td><td>Isolated shells</td><td>Freshwater snail of the family <a href="/facts/Provalvatidae/VR6mhSyR">Provalvatidae</a></td><td></td></tr><tr><td><a href="/facts/Valvatidae/hEM59nde">Valvatidae</a><a class="footnote-ref" id="fnref:27" href="#fn:27"><sup>27</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Isolated shells</td><td>Freshwater snails</td><td></td></tr></tbody></table> <h3>Fish</h3> <p>Abundant osteichthyan ichthyoliths are know. An isolated spine displaying characters seen in extant <a href="/facts/Cypriniformes/U5H2jpsr">cypriniform</a> or <a href="/facts/Siluriform/SHrSxpuH">siluriform</a> fishes has been recovered, alternatively can belong to a member of <a href="/facts/Myriacanthidae/SHazTmyG">Myriacanthidae</a>.<a class="footnote-ref" id="fnref:28" href="#fn:28"><sup>28</sup></a> </p> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Arganodus/raNutt9z">Arganodus</a></i><a class="footnote-ref" id="fnref:29" href="#fn:29"><sup>29</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Isolated tooth plates</td><td><a href="/facts/Lungfish/oNNrFFoE">Lungfish</a></td><td></td></tr><tr><td><i><a href="/facts/Lepidotes/E4MJVMhb">Lepidotes</a></i>/<i><a href="/facts/Scheenstia/hGF04Ddm">Scheenstia</a><a class="footnote-ref" id="fnref:30" href="#fn:30"><sup>30</sup></a></i></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Isolated teeth and scales</td><td><a href="/facts/Lepidotidae/IC60Imuj">Lepidotidae</a> bony fish</td><td></td></tr><tr><td>cf. Ionoscopiformes<a class="footnote-ref" id="fnref:31" href="#fn:31"><sup>31</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Rhomboidal scales</td><td>Tentatively assigned to Ophiopsidae</td><td></td></tr><tr><td><a href="/facts/Mawsoniidae/LhqA6Guz">Mawsoniidae</a><a class="footnote-ref" id="fnref:32" href="#fn:32"><sup>32</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Skull bones including a parasphenoid</td><td><a href="/facts/Coelacanth/TaEXps8Q">Coelacanth</a></td><td></td></tr><tr><td><a href="/facts/Osteoglossiformes/DekXqN05">Osteoglossiformes</a><a class="footnote-ref" id="fnref:33" href="#fn:33"><sup>33</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Squamules (small scales)</td><td>The oldest record of the group</td><td></td></tr></tbody></table> <h3>Amphibians</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><a href="/facts/Albanerpetontidae/sCRewqvg">Albanerpetontidae</a><a class="footnote-ref" id="fnref:34" href="#fn:34"><sup>34</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Incomplete premaxilla and incomplete frontal</td><td>Likely represents a new taxon</td><td></td></tr><tr><td><a href="/facts/Caudata/pCeXrHka">Caudata</a>?<a class="footnote-ref" id="fnref:35" href="#fn:35"><sup>35</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Fragment of a dentary</td><td></td><td></td></tr><tr><td><a href="/facts/Lissamphibia/zeV5XtE9">Lissamphibia</a>?<a class="footnote-ref" id="fnref:36" href="#fn:36"><sup>36</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Fragment of a maxilla</td><td></td><td></td></tr></tbody></table> <h3>Turtles</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><a href="/facts/Araripemydidae/eUVXtacL">Araripemydidae</a>?<a class="footnote-ref" id="fnref:37" href="#fn:37"><sup>37</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Shell Fragments</td><td>Resembles both <a href="/facts/Araripemydidae/eUVXtacL">Araripemydids</a> and <a href="/facts/Trionychidae/LKKs8MY3">Trionychidae</a></td><td></td></tr><tr><td><a href="/facts/Cryptodira/F85cj55N">Cryptodira</a>?<a class="footnote-ref" id="fnref:38" href="#fn:38"><sup>38</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Shell Fragments</td><td>Resembles younger-aged <a href="/facts/Solemydidae/pmemL38s">Solemydidae</a> and <a href="/facts/Pleurosternidae/8NYCnCVQ">Pleurosternidae</a> taxa</td><td></td></tr><tr><td><a href="/facts/Testudinata/R7d1PlXs">Testudinata</a><a class="footnote-ref" id="fnref:39" href="#fn:39"><sup>39</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Shell Fragments</td><td>Neural plate slightly longer than wide</td><td></td></tr><tr><td><a href="/facts/Testudinata/R7d1PlXs">Testudinata</a><a class="footnote-ref" id="fnref:40" href="#fn:40"><sup>40</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Shell Fragments</td><td>Represented by smooth plates</td><td></td></tr></tbody></table> <h3>Lepidosaurs</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Parviraptor/ov0hTqFa">Parviraptor</a><a class="footnote-ref" id="fnref:41" href="#fn:41"><sup>41</sup></a></i></td><td><i>Cf.P.sp.</i></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Four vertebral centra</td><td><a href="/facts/Squamata/sM1oc2vj">Squamatan</a>, previously identified as a stem-snake</td><td></td></tr><tr><td><a href="/facts/Rhynchocephalia/WftNntW9">Rhynchocephalia</a><a class="footnote-ref" id="fnref:42" href="#fn:42"><sup>42</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>One fragment of dentary bearing two incomplete teeth, one fragment of maxilla bearing the bases of two teeth.</td><td>Similar to the co-regional but younger genus <i><a href="/facts/Tingitana/omtPHASK">Tingitana</a></i>, but also to <i><a href="/facts/Clevosaurus/e7PbKatS">Clevosaurus</a></i>.</td><td></td></tr><tr><td><a href="/facts/Scincomorpha/7KAKtbLM">Scincomorpha</a><a class="footnote-ref" id="fnref:43" href="#fn:43"><sup>43</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>A fragment of bone bearing teeth, potentially a fragment of bone bearing one complete tooth</td><td><a href="/facts/Squamata/sM1oc2vj">Squamatan</a></td><td></td></tr><tr><td><a href="/facts/Squamata/sM1oc2vj">Squamata</a><a class="footnote-ref" id="fnref:44" href="#fn:44"><sup>44</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Incomplete maxilla, incomplete dorsal vertebra, one proximal part of femur</td><td></td><td></td></tr></tbody></table> <h3>Choristoderes</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><a href="/facts/Choristodera/rTolEzdL">Choristodera</a><a class="footnote-ref" id="fnref:45" href="#fn:45"><sup>45</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Fragmentary dentary, centrum of an anterior caudal vertebra, possibly centrum of a dorsal vertebra</td><td>Similar to <i><a href="/facts/Cteniogenys/MbKeYLcf">Cteniogenys</a></i></td><td></td></tr></tbody></table> <h3>Dinosaurs</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><a href="/facts/Ornithischia/g1SAFJvB">Ornithischia</a><a class="footnote-ref" id="fnref:46" href="#fn:46"><sup>46</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Tooth</td><td>Similar to <i><a href="/facts/Alocodon/fJ7o649S">Alocodon</a></i></td><td></td></tr><tr><td>cf.<a href="/facts/Stegosauria/vLTt2MS1">Stegosauria</a><a class="footnote-ref" id="fnref:47" href="#fn:47"><sup>47</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Poorly preserved tooth</td><td></td><td></td></tr><tr><td><a href="/facts/Theropoda/KUzy1Ef7">Theropoda</a><a class="footnote-ref" id="fnref:48" href="#fn:48"><sup>48</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Several teeth</td><td></td><td></td></tr></tbody></table> <h3>Pterosaurs</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><a href="/facts/Pterosaur/YNQwa1ru">Pterosauria</a><a class="footnote-ref" id="fnref:49" href="#fn:49"><sup>49</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Teeth</td><td>Similar to <a href="/facts/Wukongopteridae/P55TYcXB">wukongopterids</a></td><td></td></tr><tr><td><a href="/facts/Rhamphorhynchidae/SApqVJjn">Rhamphorhynchidae</a><a class="footnote-ref" id="fnref:50" href="#fn:50"><sup>50</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Teeth</td><td></td><td></td></tr></tbody></table> <h3>Crocodyliformes</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><a href="/facts/Atoposauridae/11RHcT4A">Atoposauridae</a><a class="footnote-ref" id="fnref:51" href="#fn:51"><sup>51</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Teeth</td><td></td><td></td></tr><tr><td><a href="/facts/Teleosauridae/rMecm599">Teleosauridae</a><a class="footnote-ref" id="fnref:52" href="#fn:52"><sup>52</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Teeth</td><td></td><td></td></tr><tr><td><a href="/facts/Thalattosuchia/G4CsVybF">Thalattosuchia</a><a class="footnote-ref" id="fnref:53" href="#fn:53"><sup>53</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Teeth, various skeletal elements probably belonging to a single individual</td><td></td><td></td></tr><tr><td><i><a href="/facts/Theriosuchus/BcpkJwQB">Theriosuchus</a><a class="footnote-ref" id="fnref:54" href="#fn:54"><sup>54</sup></a></i></td><td><i>Cf.T. sp.</i></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Teeth</td><td>An <a href="/facts/Atoposauridae/11RHcT4A">Atoposaurid</a></td><td></td></tr></tbody></table> <h3>Mammals</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><a href="/facts/Amphitheriida/E5X3AHYs">Amphitheriidae</a><a class="footnote-ref" id="fnref:55" href="#fn:55"><sup>55</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Fragment of right dentary of a juvenile individual with an anterior lower premolar and a replacing posterior premolar, fragment of an edentulous right dentary, probably middle-posterior part.</td><td>Both fragments probably belong to the same taxon</td><td></td></tr><tr><td><a href="/facts/Dryolestida/BXQXpTsN">Dryolestida</a><a class="footnote-ref" id="fnref:56" href="#fn:56"><sup>56</sup></a></td><td>Indeterminate</td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Fragment of a tooth bearing one cusp and one root</td><td></td><td></td></tr></tbody></table> <h3>Charophyta</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i>Aclistochara</i><a class="footnote-ref" id="fnref:57" href="#fn:57"><sup>57</sup></a></td><td><i>A. africana</i></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Gyrogonites</td><td>A Characean algae of the family <a href="/facts/Characeae/rr9rLEbV">Characeae</a></td><td></td></tr><tr><td><i>Porochara<a class="footnote-ref" id="fnref:58" href="#fn:58"><sup>58</sup></a></i></td><td><i>P. hians</i></td><td>Anoual</td><td>Lower Member</td><td><a href="/facts/Oogonia/cj3muKlR">Oogonia</a></td><td>A characean algae of the family Porocharaceae</td><td></td></tr></tbody></table> <h3>Palynology</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i>Alisporites<a class="footnote-ref" id="fnref:59" href="#fn:59"><sup>59</sup></a></i></td><td><ul><li><i>A.</i> spp.</li></ul></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Pollen</td><td>Affinities with the families <a href="/facts/Peltaspermaceae/YbKvx8UJ">Peltaspermaceae</a> or <a href="/facts/Corystospermaceae/6jeSswxr">Corystospermaceae</a></td><td></td></tr><tr><td><i>Araucariacites<a class="footnote-ref" id="fnref:60" href="#fn:60"><sup>60</sup></a></i></td><td><ul><li><i>A. australis</i></li></ul></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Pollen</td><td>Affinities with <a href="/facts/Araucariaceae/Rn6BKUbo">Araucariaceae</a> inside <a href="/facts/Coniferae/vDUgqxCx">Coniferae</a>.</td><td rowspan="2"></td></tr><tr><td><i>Callialasporites<a class="footnote-ref" id="fnref:61" href="#fn:61"><sup>61</sup></a></i></td><td><ul><li><i>C.</i> spp.</li></ul></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Pollen</td><td>Affinities with <a href="/facts/Araucariaceae/Rn6BKUbo">Araucariaceae</a> inside <a href="/facts/Coniferae/vDUgqxCx">Coniferae</a>.</td></tr><tr><td><i><a href="/facts/Classopollis/ey1lqNly">Classopollis</a><a class="footnote-ref" id="fnref:62" href="#fn:62"><sup>62</sup></a></i></td><td><ul><li><i>C.</i> spp.</li></ul></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Pollen</td><td>Affinities with <a href="/facts/Cheirolepidiaceae/ey1lqNly">Cheirolepidiaceae</a> inside <a href="/facts/Coniferae/vDUgqxCx">Coniferae</a>.</td><td></td></tr><tr><td><i>Cyathidites<a class="footnote-ref" id="fnref:63" href="#fn:63"><sup>63</sup></a></i></td><td><ul><li><i>C.</i> sp.</li></ul></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Spores</td><td>Affinities with the family <a href="/facts/Cyatheaceae/LTEajp4W">Cyatheaceae</a> inside <a href="/facts/Cyatheales/gaYAIp64">Cyatheales</a>. Arboreal Fern Spores</td><td></td></tr><tr><td><i>Cycadopites<a class="footnote-ref" id="fnref:64" href="#fn:64"><sup>64</sup></a></i></td><td><ul><li><i>C.</i> spp.</li></ul></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Pollen</td><td>Affinities with the <a href="/facts/Cycadopsida/Kc2JDExr">Cycadopsida</a> inside <a href="/facts/Cycadales/Kc2JDExr">Cycadales</a>. Pollen related with modern Cycas</td><td></td></tr><tr><td><i>Inaperturopollenites<a class="footnote-ref" id="fnref:65" href="#fn:65"><sup>65</sup></a></i></td><td><ul><li><i>I. spp.</i></li></ul></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Pollen</td><td>Affinities with the <a href="/facts/Pinidae/UnBqB49j">Pinidae</a> inside <a href="/facts/Coniferae/vDUgqxCx">Coniferae</a>.</td><td></td></tr><tr><td><i>Ischyosporites<a class="footnote-ref" id="fnref:66" href="#fn:66"><sup>66</sup></a></i></td><td><ul><li><i>I.</i> cf. <i>variegatus</i></li><li><i>I.</i> sp</li></ul></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Spores</td><td>Affinities with <a href="/facts/Pteridopsida/FV2SSw5j">Pteridopsida</a>, likely derived from a member of <a href="/facts/Anemiaceae/U2r4Pw2c">Anemiaceae</a></td><td></td></tr><tr><td><i>Leptolepidites<a class="footnote-ref" id="fnref:67" href="#fn:67"><sup>67</sup></a></i></td><td><ul><li><i>L. spp.</i></li></ul></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Spores</td><td>Affinities with the family <a href="/facts/Dennstaedtiaceae/brLj1myo">Dennstaedtiaceae</a> in the <a href="/facts/Polypodiales/aCUmdcLK">Polypodiales</a>. Forest fern spores.</td><td></td></tr><tr><td><i>Pityosporites<a class="footnote-ref" id="fnref:68" href="#fn:68"><sup>68</sup></a></i></td><td><ul><li><i>P. sp.</i></li></ul></td><td>Guelb el Ahmar</td><td>Lower Member</td><td>Pollen</td><td>Affinities with the <a href="/facts/Pinidae/UnBqB49j">Pinidae</a> inside <a href="/facts/Coniferae/vDUgqxCx">Coniferae</a>.</td><td></td></tr></tbody></table> <h3>Fossil Wood</h3> <table><tbody><tr><th>Genus</th><th>Species</th><th>Location</th><th>Stratigraphic Position</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Agathoxylon/WzwtrbYP">Agathoxylon</a></i><a class="footnote-ref" id="fnref:69" href="#fn:69"><sup>69</sup></a></td><td><ul><li><i>A.</i> aff. <i>sahariense</i></li></ul></td><td><ul><li>Guelb el Ahmar</li><li>Oued Metlili</li></ul></td><td>Lower Member</td><td>Fossil Wood</td><td>Conifer fossil wood of the family <a href="/facts/Araucariaceae/Rn6BKUbo">Araucariaceae</a></td><td></td></tr><tr><td><i><a href="/facts/Metapodocarpoxylon/BJArC2RL">Metapodocarpoxylon</a></i><a class="footnote-ref" id="fnref:70" href="#fn:70"><sup>70</sup></a></td><td><ul><li><i>M. libanoticum</i></li></ul></td><td><ul><li>Guelb el Ahmar</li><li>Oued Metlili</li></ul></td><td>Lower Member</td><td>Fossil Wood</td><td>Conifer fossil wood of the family <a href="/facts/Podocarpaceae/C89JGX6w">Podocarpaceae</a></td><td></td></tr><tr><td><i>Protopodocarpoxylon</i><a class="footnote-ref" id="fnref:71" href="#fn:71"><sup>71</sup></a></td><td><ul><li><i>P. subrochii</i></li><li><i>P. teixeirae</i></li></ul></td><td><ul><li>Guelb el Ahmar</li><li>Oued Metlili</li></ul></td><td>Lower Member</td><td>Fossil Wood</td><td>Conifer fossil wood of the family <a href="/facts/Hirmeriellaceae/ey1lqNly">Hirmeriellaceae</a></td><td></td></tr><tr><td><i>Phyllocladoxylon</i><a class="footnote-ref" id="fnref:72" href="#fn:72"><sup>72</sup></a></td><td><ul><li><i>P. atlasicum</i></li></ul></td><td><ul><li>Guelb el Ahmar</li><li>Oued Metlili</li></ul></td><td>Lower Member</td><td>Fossil Wood</td><td>Conifer fossil wood of the family <a href="/facts/Podocarpaceae/C89JGX6w">Podocarpaceae</a></td><td></td></tr></tbody></table> <h2 id="see-also">See Also</h2> <ul><li><a href="/facts/Ksar_Metlili_Formation/QRU8Xzfv">Ksar Metlili Formation</a>, from the Same area</li> <li><a href="/facts/Guettioua_Formation/PEQ8Wf0K">Guettioua Formation</a>, coeval from the Azilal Region</li> <li><a href="/facts/El_Mers_Group/Ayc9K2WW">El Mers Group</a>, coeval from the Skoura syncline</li> <li><a href="/facts/Tafraout_Group/Fmpn84QC">Tafraout Group</a>, older, from the Central High Atlas</li> <li><a href="/facts/Azilal_Formation/L8I6dJ9e">Azilal Formation</a>, older, from the Central High Atlas</li></ul> <h2 id="references">References</h2> <ol> <li id="fn:1"><p>Haddoumi, Hamid; Charrière, André; Andreu, Bernard; Mojon, Pierre-Olivier (2008). "Les dépôts continentaux du Jurassique moyen au Crétacé inférieur dans le Haut Atlas oriental (Maroc) : paléoenvironnements successifs et signification paléogéographique". Carnets de Géologie / Notebooks on Geology. CG2008 (A06): 1–29. <a href="https://hal.science/hal-00308920/" target="_blank">https://hal.science/hal-00308920/</a> <a href="#fnref:1" class="footnote-back-ref">↩</a></p></li> <li id="fn:2"><p>Haddoumi, Hamid; Allain, Ronan; Meslouh, Said; Metais, Grégoire; Monbaron, Michel; Pons, Denise; Rage, Jean-Claude; Vullo, Romain; Zouhri, Samir (January 2016). "Guelb el Ahmar (Bathonian, Anoual Syncline, eastern Morocco): First continental flora and fauna including mammals from the Middle Jurassic of Africa" (PDF). Gondwana Research. 29 (1): 290–319. Bibcode:2016GondR..29..290H. doi:10.1016/j.gr.2014.12.004. ISSN 1342-937X. <a href="http://doc.rero.ch/record/258861/files/mon_gab.pdf" target="_blank">http://doc.rero.ch/record/258861/files/mon_gab.pdf</a> <a href="#fnref:2" class="footnote-back-ref">↩</a></p></li> <li id="fn:3"><p>Haddoumi, Hamid; Aiméras, Yves; Bodergat, Anne-Marie; Charrière, André; Mangold, Charles; Benshili, Khadija (1998-07-01). "Âges et environnements des Couches rouges d'Anoual (Jurassique moyen et Crétacé inférieur, Haut-Atlas oriental, Maroc)". Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science. 327 (2): 127–133. doi:10.1016/S1251-8050(98)80043-8. ISSN 1251-8050. <a href="https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438" target="_blank">https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438</a> <a href="#fnref:3" class="footnote-back-ref">↩</a></p></li> <li id="fn:4"><p>Haddoumi, Hamid; Aiméras, Yves; Bodergat, Anne-Marie; Charrière, André; Mangold, Charles; Benshili, Khadija (1998-07-01). "Âges et environnements des Couches rouges d'Anoual (Jurassique moyen et Crétacé inférieur, Haut-Atlas oriental, Maroc)". Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science. 327 (2): 127–133. doi:10.1016/S1251-8050(98)80043-8. ISSN 1251-8050. <a href="https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438" target="_blank">https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438</a> <a href="#fnref:4" class="footnote-back-ref">↩</a></p></li> <li id="fn:5"><p>Haddoumi, Hamid; Aiméras, Yves; Bodergat, Anne-Marie; Charrière, André; Mangold, Charles; Benshili, Khadija (1998-07-01). "Âges et environnements des Couches rouges d'Anoual (Jurassique moyen et Crétacé inférieur, Haut-Atlas oriental, Maroc)". Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science. 327 (2): 127–133. doi:10.1016/S1251-8050(98)80043-8. ISSN 1251-8050. <a href="https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438" target="_blank">https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438</a> <a href="#fnref:5" class="footnote-back-ref">↩</a></p></li> <li id="fn:6"><p>Haddoumi, Hamid; Aiméras, Yves; Bodergat, Anne-Marie; Charrière, André; Mangold, Charles; Benshili, Khadija (1998-07-01). "Âges et environnements des Couches rouges d'Anoual (Jurassique moyen et Crétacé inférieur, Haut-Atlas oriental, Maroc)". Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science. 327 (2): 127–133. doi:10.1016/S1251-8050(98)80043-8. ISSN 1251-8050. <a href="https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438" target="_blank">https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438</a> <a href="#fnref:6" class="footnote-back-ref">↩</a></p></li> <li id="fn:7"><p>Haddoumi, Hamid; Aiméras, Yves; Bodergat, Anne-Marie; Charrière, André; Mangold, Charles; Benshili, Khadija (1998-07-01). "Âges et environnements des Couches rouges d'Anoual (Jurassique moyen et Crétacé inférieur, Haut-Atlas oriental, Maroc)". Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science. 327 (2): 127–133. doi:10.1016/S1251-8050(98)80043-8. ISSN 1251-8050. <a href="https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438" target="_blank">https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438</a> <a href="#fnref:7" class="footnote-back-ref">↩</a></p></li> <li id="fn:8"><p>Haddoumi, Hamid; Aiméras, Yves; Bodergat, Anne-Marie; Charrière, André; Mangold, Charles; Benshili, Khadija (1998-07-01). "Âges et environnements des Couches rouges d'Anoual (Jurassique moyen et Crétacé inférieur, Haut-Atlas oriental, Maroc)". Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science. 327 (2): 127–133. doi:10.1016/S1251-8050(98)80043-8. ISSN 1251-8050. <a href="https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438" target="_blank">https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438</a> <a href="#fnref:8" class="footnote-back-ref">↩</a></p></li> <li id="fn:9"><p>Haddoumi, Hamid; Aiméras, Yves; Bodergat, Anne-Marie; Charrière, André; Mangold, Charles; Benshili, Khadija (1998-07-01). "Âges et environnements des Couches rouges d'Anoual (Jurassique moyen et Crétacé inférieur, Haut-Atlas oriental, Maroc)". Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science. 327 (2): 127–133. doi:10.1016/S1251-8050(98)80043-8. 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ISSN 1251-8050. <a href="https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438" target="_blank">https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438</a> <a href="#fnref:10" class="footnote-back-ref">↩</a></p></li> <li id="fn:11"><p>Haddoumi, Hamid; Aiméras, Yves; Bodergat, Anne-Marie; Charrière, André; Mangold, Charles; Benshili, Khadija (1998-07-01). "Âges et environnements des Couches rouges d'Anoual (Jurassique moyen et Crétacé inférieur, Haut-Atlas oriental, Maroc)". Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science. 327 (2): 127–133. doi:10.1016/S1251-8050(98)80043-8. ISSN 1251-8050. <a href="https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438" target="_blank">https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438</a> <a href="#fnref:11" class="footnote-back-ref">↩</a></p></li> <li id="fn:12"><p>Haddoumi, Hamid; Aiméras, Yves; Bodergat, Anne-Marie; Charrière, André; Mangold, Charles; Benshili, Khadija (1998-07-01). "Âges et environnements des Couches rouges d'Anoual (Jurassique moyen et Crétacé inférieur, Haut-Atlas oriental, Maroc)". Comptes Rendus de l'Académie des Sciences - Series IIA - Earth and Planetary Science. 327 (2): 127–133. doi:10.1016/S1251-8050(98)80043-8. ISSN 1251-8050. <a href="https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438" target="_blank">https://www.sciencedirect.com/science/article/abs/pii/S1251805098800438</a> <a href="#fnref:12" class="footnote-back-ref">↩</a></p></li> <li id="fn:13"><p>Haddoumi, Hamid; Charrière, André; Andreu, Bernard; Mojon, Pierre-Olivier (2008). "Les dépôts continentaux du Jurassique moyen au Crétacé inférieur dans le Haut Atlas oriental (Maroc) : paléoenvironnements successifs et signification paléogéographique". Carnets de Géologie / Notebooks on Geology. CG2008 (A06): 1–29. <a href="https://hal.science/hal-00308920/" target="_blank">https://hal.science/hal-00308920/</a> <a href="#fnref:13" class="footnote-back-ref">↩</a></p></li> <li id="fn:14"><p>Haddoumi, Hamid; Charrière, André; Andreu, Bernard; Mojon, Pierre-Olivier (2008). 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