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Anacleto Formation
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<h2 id="fossil-content">Fossil content</h2> <table><tbody><tr><th></th><th></th><th></th></tr><tr><td>Color key<table><tbody><tr><td><a href="/facts/Taxon/caQC1e87">Taxon</a></td><td>Reclassified taxon</td><td>Taxon falsely reported as present</td><td>Dubious taxon or junior synonym</td><td><a href="/facts/Ichnotaxon/edWeWT20">Ichnotaxon</a></td><td><a href="/facts/Ootaxon/nH5XHNRS">Ootaxon</a></td><td><a href="/facts/Morphotaxon/zWsaRMUe">Morphotaxon</a></td></tr></tbody></table></td><td></td><td>Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.</td></tr></tbody></table> <p>The following animals are known from bones found in the Anacleto Formation: </p> <ul><li>several species of <a href="/facts/Lizard/VPCpCwjT">lizards</a></li> <li>several species of <a href="/facts/Mammal/phSwTzBo">mammals</a></li></ul> <h3>Crocodylomorphs</h3> <table><tbody><tr><th colspan="6"><a href="/facts/Crocodylomorph/clPyHg6D">Crocodylomorphs</a> reported from the Anacleto Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Presence</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Gasparinisuchus/U5hma5dE">Gasparinisuchus</a></i></td><td><i>G. peirosauroides</i></td><td>Cañadón Amarrillo, south <a href="/facts/Malarg%C3%BCe/qofC6ToY">Malargüe</a> city, <a href="/facts/Mendoza_Province/p8q23PAs">Mendoza Province</a>.<a class="footnote-ref" id="fnref:4" href="#fn:4"><sup>4</sup></a></td><td>PV-CRIDC-12 (right <a href="/facts/Premaxilla/82BL4FgY">premaxilla</a> and <a href="/facts/Maxilla/LbgmySaT">maxilla</a>, isolated teeth, and part of the postcranial skeleton).<a class="footnote-ref" id="fnref:5" href="#fn:5"><sup>5</sup></a></td><td>A <a href="/facts/Peirosauridae/s307kNA2">peirosaurid</a> also known from the <a href="/facts/Bajo_de_la_Carpa_Formation/NdS5UEkN">Bajo de la Carpa Formation</a>.</td><td></td></tr><tr><td><i><a href="/facts/Peirosaurus/aVFLyedp">Peirosaurus</a></i></td><td><i>P. torminni</i></td><td>Cañadón Amarrillo, south <a href="/facts/Malarg%C3%BCe/qofC6ToY">Malargüe</a> city, <a href="/facts/Mendoza_Province/p8q23PAs">Mendoza Province</a>.<a class="footnote-ref" id="fnref:6" href="#fn:6"><sup>6</sup></a></td><td>PV-CRIDC-12 (right <a href="/facts/Premaxilla/82BL4FgY">premaxilla</a> and <a href="/facts/Maxilla/LbgmySaT">maxilla</a>, isolated teeth, and part of the postcranial skeleton).<a class="footnote-ref" id="fnref:7" href="#fn:7"><sup>7</sup></a></td><td>Specimen reassigned to <i><a href="/facts/Gasparinisuchus/U5hma5dE">Gasparinisuchus</a></i>.</td><td></td></tr></tbody></table> <h3>Dinosaurs</h3> <h4>Ornithischians</h4> <table><tbody><tr><th colspan="7"><a href="/facts/Ornithischia/g1SAFJvB">Ornithischians</a> reported from the Anacleto Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Stratigraphic position</th><th>Presence</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Gasparinisaura/FXWlCe6d">Gasparinisaura</a></i></td><td><i>G. cincosaltensis</i></td><td></td><td>Cinco Saltos, <a href="/facts/R%C3%ADo_Negro_Province/4Ha1j8Be">Río Negro Province</a>.<a class="footnote-ref" id="fnref:8" href="#fn:8"><sup>8</sup></a></td><td>Multiple specimens.<a class="footnote-ref" id="fnref:9" href="#fn:9"><sup>9</sup></a></td><td>A small <a href="/facts/Ornithopoda/HpWlQsXI">ornithopod</a>.</td><td></td></tr></tbody></table> <h4>Sauropods</h4> <p><a href="/facts/Nest/7tq5lt4V">Nests</a> of <a href="/facts/Dinosaur_egg/bBypgTYQ">dinosaur eggs</a>, many with preserved <a href="/facts/Embryo/9QHnLVLN">embryos</a> inside, have been discovered in large quantities at the famous <a href="/facts/Auca_Mahuevo/UwGGyfHa">Auca Mahuevo</a> locality, and have been attributed to titanosaurs.<a class="footnote-ref" id="fnref:10" href="#fn:10"><sup>10</sup></a> </p> <table><tbody><tr><th colspan="7"><a href="/facts/Sauropod/8Iqh2yEJ">Sauropods</a> reported from the Anacleto Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Stratigraphic Position</th><th>Presence</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Antarctosaurus/Q2xBWWbq">Antarctosaurus</a></i></td><td><i>A. wichmannianus</i></td><td></td><td><a class="footnote-ref" id="fnref:11" href="#fn:11"><sup>11</sup></a></td><td></td><td>A <a href="/facts/Titanosauria/QzkgrcQR">titanosaur</a>.</td><td></td></tr><tr><td><i><a href="/facts/Barrosasaurus/XbLkeDu8">Barrosasaurus</a></i></td><td><i>B. casamiquelai</i></td><td>Lower</td><td><a href="/facts/Neuqu%C3%A9n_Province/RaQYfAIJ">Neuquén Province</a>.<a class="footnote-ref" id="fnref:12" href="#fn:12"><sup>12</sup></a></td><td>Vertebrae.<a class="footnote-ref" id="fnref:13" href="#fn:13"><sup>13</sup></a></td><td>A <a href="/facts/Titanosauria/QzkgrcQR">titanosaur</a>.</td><td></td></tr><tr><td><i><a href="/facts/Chadititan/HMbRYrHn">Chadititan</a></i><a class="footnote-ref" id="fnref:14" href="#fn:14"><sup>14</sup></a></td><td><i>C. calvoi</i></td><td>Lower</td><td>General Roca city, Río Negro Province.</td><td>A few fragment holotype consists of caudal vertebrae, partial humeri, femur bone, ulna, and pubis.</td><td>A <a href="/facts/Titanosauria/QzkgrcQR">titanosaur</a>.</td><td></td></tr><tr><td><i><a href="/facts/Laplatasaurus/bzrKP4Xk">Laplatasaurus</a></i></td><td><i>L. araukanicus</i></td><td>Uppermost</td><td>Several localities in Patagonia.<a class="footnote-ref" id="fnref:15" href="#fn:15"><sup>15</sup></a></td><td>Multiple specimens.<a class="footnote-ref" id="fnref:16" href="#fn:16"><sup>16</sup></a></td><td>A <a href="/facts/Titanosauria/QzkgrcQR">titanosaur</a>.</td><td></td></tr><tr><td><i><a href="/facts/Narambuenatitan/hTYLp24J">Narambuenatitan</a></i></td><td><i>N. palomoi</i></td><td>Lower</td><td></td><td>Remains of a subadult.<a class="footnote-ref" id="fnref:17" href="#fn:17"><sup>17</sup></a></td><td>A <a href="/facts/Titanosauria/QzkgrcQR">titanosaur</a>.</td><td></td></tr><tr><td><i><a href="/facts/Neuquensaurus/z5glB0bp">Neuquensaurus</a></i></td><td><i>N. australis</i></td><td>Uppermost</td><td>Cinco Saltos, <a href="/facts/R%C3%ADo_Negro_Province/4Ha1j8Be">Río Negro Province</a>.<a class="footnote-ref" id="fnref:18" href="#fn:18"><sup>18</sup></a></td><td>Six caudal vertebrae.</td><td>A <a href="/facts/Saltasaurinae/gosvgGDB">saltasaurine</a>.</td><td></td></tr><tr><td><i><a href="/facts/Pitekunsaurus/pRcvdI2B">Pitekunsaurus</a></i></td><td><i>P. macayai</i></td><td></td><td></td><td>"Braincase, left frontal, one tooth, four <a href="/facts/Cervical_vertebrae/M7Kf14h9">cervical vertebrae</a>, three dorsal vertebrae, four caudal vertebrae, right ulna and scapula, proximal extreme of left <a href="/facts/Femur/uVyTwj4l">femur</a>, rib fragments and uncertain remains".<a class="footnote-ref" id="fnref:19" href="#fn:19"><sup>19</sup></a></td><td>A <a href="/facts/Titanosauria/QzkgrcQR">titanosaur</a>.</td><td></td></tr><tr><td><i><a href="/facts/Teratopodus/lGn0PqnL">Teratopodus</a></i></td><td><i>T. malarguensis</i></td><td></td><td>Southern <a href="/facts/Mendoza_Province/p8q23PAs">Mendoza Province</a>.<a class="footnote-ref" id="fnref:20" href="#fn:20"><sup>20</sup></a></td><td>Footprint trackways.<a class="footnote-ref" id="fnref:21" href="#fn:21"><sup>21</sup></a></td><td>A <a href="/facts/Titanosauria/QzkgrcQR">titanosaur</a> <a href="/facts/Ichnotaxon/edWeWT20">ichnotaxon</a>.</td><td></td></tr></tbody></table> <h4>Theropods</h4> <p>The oldest known unequivocal <a href="/facts/Bird/RJF6zwLP">bird</a> footprints from <a href="/facts/South_America/PhrFEMk7">South America</a> were discovered in the Anacleto Formation. The small footprints were tentatively assigned to the <a href="/facts/Ichnogenus/edWeWT20">ichnogenus</a> <i>Aquatilavipes</i> and might have been produced by <i><a href="/facts/Patagopteryx/qq7g27l0">Patagopteryx</a></i> (whose fossils were only found in the Bajo de la Carpa Formation however) or some unknown <a href="/facts/Wader/ncAbzR2k">wader</a>-like bird; they lack a hind toe. <i><a href="/facts/Ignotornis/1SR9thdb">Ignotornis</a></i> refers to similar footprints made by larger birds with a small hind toe; they might have been left by <i><a href="/facts/Neuquenornis/RcgDJtF7">Neuquenornis</a></i>, but this is also only known from the Bajo de la Carpa Formation. Footprints of these two ichnogenera have also been found elsewhere, but it must be understood that assignment to the same ichnogenus does not imply a close relatedness of the organisms that produced these traces, only a similar <a href="/facts/Morphology_(biology)/Nl3B0myy">morphology</a>.<a class="footnote-ref" id="fnref:22" href="#fn:22"><sup>22</sup></a> </p><p>Even smaller and somewhat unusual footprints assigned to <i>Barrosopus</i> are only known from the Anacleto Formation. They were almost certainly made by some tiny theropod, but whether this was a bird is not quite clear: the innermost front toes of the animal leaving these tracks attached in a position higher than the others. In that, and in their dimensions, they are a very close match for the odd-footed <a href="/facts/Enantiornithine/kuC8WyFS">enantiornithine</a> bird <i><a href="/facts/Yungavolucris_brevipedalis/6x7EhRsI">Yungavolucris brevipedalis</a></i>, but this is only known from the <a href="/facts/Maastrichtian/xRBafamu">Maastrichtian</a> <a href="/facts/Lecho_Formation/gjQnbT87">Lecho Formation</a> which is some 10 million years younger.<a class="footnote-ref" id="fnref:23" href="#fn:23"><sup>23</sup></a> </p> <table><tbody><tr><th colspan="7"><a href="/facts/Theropod/KUzy1Ef7">Theropods</a> reported from the Anacleto Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Stratigraphic Position</th><th>Presence</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><a href="/facts/Abelisauridae/nvdLVChd">Abelisauridae</a></td><td>Indeterminate</td><td>Lower</td><td>Northwest Patagonia.<a class="footnote-ref" id="fnref:24" href="#fn:24"><sup>24</sup></a><a class="footnote-ref" id="fnref:25" href="#fn:25"><sup>25</sup></a></td><td>MPCN-PV 69, consisting of a partial <a href="/facts/Premaxilla/82BL4FgY">premaxilla</a>, fragmentary vertebrae, proximal portion of both <a href="/facts/Humerus/9h0R2GuY">humeri</a>, distal portion of the <a href="/facts/Pubis_(bone)/4gU0dgIy">pubis</a>, and an incomplete pedal ungual.<a class="footnote-ref" id="fnref:26" href="#fn:26"><sup>26</sup></a><a class="footnote-ref" id="fnref:27" href="#fn:27"><sup>27</sup></a></td><td>An indeterminate <a href="/facts/Abelisauridae/nvdLVChd">abelisaurid</a>.</td><td></td></tr><tr><td><i><a href="/facts/Abelisaurus/FV9hTWA5">Abelisaurus</a></i></td><td><i>A. comahuensis</i></td><td></td><td>Lago Pellegrini stone quarries.<a class="footnote-ref" id="fnref:28" href="#fn:28"><sup>28</sup></a></td><td>Skull.<a class="footnote-ref" id="fnref:29" href="#fn:29"><sup>29</sup></a></td><td>An <a href="/facts/Abelisauridae/nvdLVChd">abelisaurid</a>, originally thought to be from the <a href="/facts/Allen_Formation/zRhFI0qU">Allen Formation</a>.</td><td></td></tr><tr><td><i><a href="/facts/Aerosteon/jXXzc3Kc">Aerosteon</a></i></td><td><i>A. riocoloradensis</i></td><td></td><td>"Cañadon Amarillo (S 37.5°, W 70.5°), north of <a href="/facts/Cerro_Colorados/6YgLvzG3">Cerro Colorado</a>, 1 km north of the <a href="/facts/R%C3%ADo_Colorado_(Argentina)/eyk3Asen">Río Colorado</a> near the southern border of <a href="/facts/Mendoza_Province/p8q23PAs">Mendoza Province</a>, Argentina."<a class="footnote-ref" id="fnref:30" href="#fn:30"><sup>30</sup></a></td><td>Skeletal remains.<a class="footnote-ref" id="fnref:31" href="#fn:31"><sup>31</sup></a></td><td>A <a href="/facts/Megaraptoridae/lyJ9jAl3">megaraptorid</a>.</td><td></td></tr><tr><td><i><a href="/facts/Aucasaurus/vDsQf3l0">Aucasaurus</a></i></td><td><i>A. garridoi</i></td><td>Upper</td><td><a href="/facts/Auca_Mahuevo/UwGGyfHa">Auca Mahuevo</a>.<a class="footnote-ref" id="fnref:32" href="#fn:32"><sup>32</sup></a></td><td>Skeleton of an adult (MCF-PVPH-236).<a class="footnote-ref" id="fnref:33" href="#fn:33"><sup>33</sup></a></td><td>An <a href="/facts/Abelisauridae/nvdLVChd">abelisaurid</a>.</td><td></td></tr><tr><td><a href="/facts/Megaraptora/lyJ9jAl3">Megaraptora</a> indet.</td><td>Indeterminate</td><td>Upper</td><td><a href="/facts/Auca_Mahuevo/UwGGyfHa">Auca Mahuevo</a>.<a class="footnote-ref" id="fnref:34" href="#fn:34"><sup>34</sup></a></td><td>MCF-PVPH-416, a fragmentary pubic boot with unfused proximal symphysial contact.<a class="footnote-ref" id="fnref:35" href="#fn:35"><sup>35</sup></a></td><td>A large <a href="/facts/Megaraptora/lyJ9jAl3">megaraptoran</a>.</td><td></td></tr></tbody></table> <h3>Squamates</h3> <table><tbody><tr><th colspan="6"><a href="/facts/Squamates/sM1oc2vj">Squamates</a> reported from the Anacleto Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Presence</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Dinilysia/9J0HjVtQ">Dinilysia</a></i></td><td><i>D.</i> sp.</td><td>Aguada Toledo, south of <a href="/facts/Mari_Menuco_Lake/mb8y90ty">Mari Menuco Lake</a>.<a class="footnote-ref" id="fnref:36" href="#fn:36"><sup>36</sup></a></td><td>24 articulated mid-posterior trunk vertebrae with the base of their respective ribs (UNC-CIP 1).<a class="footnote-ref" id="fnref:37" href="#fn:37"><sup>37</sup></a></td><td>A large <a href="/facts/Snake/YhA2gcn0">snake</a>.</td><td></td></tr></tbody></table> <h3>Testudines</h3> <table><tbody><tr><th colspan="7"><a href="/facts/Testudines/X0ImnxYt">Testudines</a> reported from the Anacleto Formation</th></tr><tr><th>Genus</th><th>Species</th><th>Stratigraphic Position</th><th>Presence</th><th>Material</th><th>Notes</th><th>Images</th></tr><tr><td><i><a href="/facts/Prochelidella/XeRhFs3I">Prochelidella</a></i></td><td><i>P. palomoi</i></td><td></td><td>Aguada Grande site, <a href="/facts/Neuqu%C3%A9n_Province/RaQYfAIJ">Neuquén Province</a>.<a class="footnote-ref" id="fnref:38" href="#fn:38"><sup>38</sup></a></td><td>A partially preserved skull, <a href="/facts/Carapace/8VnnWH2b">carapace</a> remains, an almost complete <a href="/facts/Plastron/HRERph7C">plastron</a>, and a left <a href="/facts/Ilium_(bone)/4zEB7NX0">ilium</a> (MAU-Pv-AG-452).<a class="footnote-ref" id="fnref:39" href="#fn:39"><sup>39</sup></a></td><td>A <a href="/facts/Chelidae/dXBBdTL5">chelid</a> turtle.</td><td></td></tr><tr><td><i><a href="/facts/Yaminuechelys/MtjtLtJs">Yaminuechelys</a></i></td><td><i>Y.</i> aff. <i>maior</i></td><td>Lower</td><td><a href="/facts/Neuqu%C3%A9n_Province/RaQYfAIJ">Neuquén Province</a>.<a class="footnote-ref" id="fnref:40" href="#fn:40"><sup>40</sup></a></td><td>Two specimens (MAU-Pv-N-475 & MAU-Pv-PR-455).<a class="footnote-ref" id="fnref:41" href="#fn:41"><sup>41</sup></a></td><td>A long-necked <a href="/facts/Chelidae/dXBBdTL5">chelid</a> turtle.</td><td></td></tr></tbody></table> <h2 id="see-also">See also</h2> <ul> <li>Geology portal</li><li>Paleontology portal</li></ul> <ul><li><a href="/facts/List_of_fossil_sites/6gYhEPes">List of fossil sites</a></li> <li><a href="/facts/Los_Alamitos_Formation/MEN1sPMY">Los Alamitos Formation</a></li> <li><a href="/facts/Los_Blanquitos_Formation/HWsymNjz">Los Blanquitos Formation</a></li> <li><a href="/facts/Adamantina_Formation/hdhEMcXa">Adamantina Formation</a></li></ul> <h3>Bibliography</h3> <ul><li>Chiappe, Luis M. (1993): Enantiornithine (Aves) Tarsometatarsi from the Cretaceous Lecho Formation of Northwestern Argentina. <i>American Museum Novitates</i> 3083: 1-27. [English with Spanish abstract] <a href="http://digitallibrary.amnh.org/dspace/bitstream/2246/4981/1/N3083.pdf">PDF fulltext</a></li> <li>Coria, Rodolfo A.; Currie, Philip J.; Eberth, David & Garrido, Alberto (2002): Bird footprints from the Anacleto Formation (Late Cretaceous), Neuquén, Argentina. <i>Ameghiniana</i> 39(4): 453-463. [English with Spanish abstract] <a href="https://web.archive.org/web/20061124234955/http://www.biology.ualberta.ca/faculty/philip_currie/uploads/pdfs/2002/2002Bird_Footprints.PDF">PDF fulltext</a></li> <li>Fossa Mancini, E.; <a href="/facts/Egidio_Feruglio/XAQvwxxY">Feruglio, E.</a>; Yussen de Campana, J.C. (1938). "Una reunión de geólogos de YPF y el problema de la terminología estratigráfica ("A YPF geologists' reunion and the problem of stratigraphy terminology")". <i>Boletín de Informaciones Petroleras</i>. 15: 1–67.</li> <li>Leanza, H.A.; Apesteguia, S.; <a href="/facts/Fernando_Novas/kzH2Bzm9">Novas, F.E.</a>; De la Fuente, M.S. (2004). <a href="https://www.researchgate.net/publication/257047651">"Cretaceous terrestrial beds from the Neuquén Basin (Argentina) and their tetrapod assemblages"</a>. <i><a href="/facts/Cretaceous_Research/FBr2sFQR">Cretaceous Research</a></i>. 25 (1): 61–87. <a href="/facts/Bibcode_(identifier)/9HtdQSGB">Bibcode</a>:<a href="https://ui.adsabs.harvard.edu/abs/2004CrRes..25...61L">2004CrRes..25...61L</a>. <a href="/facts/Doi_(identifier)/muM9Etpq">doi</a>:<a href="https://doi.org/10.1016%2Fj.cretres.2003.10.005">10.1016/j.cretres.2003.10.005</a>. Retrieved 2019-02-16 – via ResearchGate.</li> <li>Lockley, Martin; Matsukawa, Masaki; Ohira, Hiroto; Li, Jianjun; Wright, Joanna; White, Diane & Chen, Peiji (2006): Bird tracks from Liaoning Province, China: New insights into avian evolution during the Jurassic-Cretaceous transition. <i>Cretaceous Research</i> 27(1): 33-43. <a href="/facts/Doi_(identifier)/muM9Etpq">doi</a>:<a href="https://doi.org/10.1016%2Fj.cretres.2005.10.007">10.1016/j.cretres.2005.10.007</a> (HTML abstract). Erratum: <a href="/facts/Doi_(identifier)/muM9Etpq">doi</a>:<a href="https://doi.org/10.1016%2Fj.cretres.2008.06.002">10.1016/j.cretres.2008.06.002</a></li> <li>Salgado, L.; Coria, R.A. & Chiappe, Luis M. (2005): Osteology of the sauropod embryos from the Upper Cretaceous of Patagonia. <i>Acta Palaeontologica Polonica</i> 50(1): 79–92. <a href="http://www.app.pan.pl/archive/published/app50/app50-079.pdf">PDF fulltext</a></li> <li>Sánchez, María Lidia; Heredia, Susana; Calvo, Jorge O. (2006). <a href="https://www.researchgate.net/publication/262615949">"Paleoambientes sedimentarios del Cretácico Superior de la Formación Plottier (Grupo Neuquén), Departamento Confluencia, Neuquén (Sedimentary paleoenvironments in the Upper Cretaceous Plottier Formation (Neuquen Group), Confluencia, Neuquén)"</a>. <i><a href="/facts/Revista_de_la_Asociaci%C3%B3n_Geol%C3%B3gica_Argentina/8SHmNSG0">Revista de la Asociación Geológica Argentina</a></i>. 61: 3–18. Retrieved 2019-02-16 – via ResearchGate.</li></ul> <h2 id="further-reading">Further reading</h2> <ul><li>S. Brizuela and A. Albino. 2011. A Scincomorpha lizard from the Campanian of Patagonia. Cretaceous Research 32:781-785</li> <li>I. A. Cerda. 2008. Gastroliths in an ornithopod dinosaur. Acta Palaeontologica Polonica 53(2):351-355</li> <li>L. M. Chiappe, R. A. Coria, L. Dingus, F. Jackson, A. Chinsamy and M. Fox. 1998. Sauropod dinosaur embryos from the Late Cretaceous of Patagonia. Nature 396:258-261</li> <li>R. A. Coria and L. M. Chiappe. 2000. Un nuevo terópodo abelisaurio de la Fm. Río Colorado (Cretácico Superior) de la Provincia del Neuquén [A new abelisaur theropod from the Río Colorado Fm. (Upper Cretaceous) of Neuquén province]. Actas XVI Jornadas Argentinas de Paleontología de Vertebrado, San Luis, Argentina 13</li> <li>L. S. Filippi and A. C. Garrido. 2012. Nuevo registro del género <i>Dinilysia</i> (Squamata, Serpentes) para la Formación Anacleto (Campaniano inferior-medio), Rincón de los Sauces, Neuquén, Argentina. Ameghiniana 49(1):132-136</li> <li>B. J. González Riga. 2011. Speeds and stance of titanosaur sauropods: analysis of <i>Titanopodus</i> tracks from the Late Cretaceous of Mendoza, Argentina. Anais da Academia Brasileira de Ciências 83(1):279-290</li> <li>A. M. Praderio, A. G. Martinelli, and C. R. A. Candeiro. 2008. Mesoeucrocodilos en el Cretácico de Malargüe: primer registro de <i>Peirosaurus tormini</i> (Crocodyliformes, Peirosauridae) para la provincia de Mendoza (Argentina). Actas del 4to. Encuentro Internacional del International Center of Earth Sciences (E-ICES-4), Malargüe, Mendoza 1-7</li> <li>L. Salgado, S. Apesteguía, and S. Heredia. 2005. A new specimen of <i>Neuquensaurus australis</i>, a Late Cretaceous saltasaurine titanosaur from North Patagonia. Journal of Vertebrate Paleontology 25(3):623-634</li> <li>L. Salgado, R. A. Coria, and S. E. Heredia. 1996. Nuevos materiales de ornitópodos (Ornithischia) en la Formación Rio Colorado (Cretácico Superior) de la Provincia de Rio Negro [New materials of ornithopods (Ornithischia) in the Rio Colorado Formation (Upper Cretaceous) of Rio Negro province]. Ameghiniana 33(4):471</li> <li>P. C. Sereno, R. N. Martínez, J. A. Wilson, D. J. Varricchio, O. A. Alcober and H. C. E. Larsson. 2008. 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