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Cow Branch Formation
Geologic formation in the eastern U.S

The Cow Branch Formation is a Late Triassic geologic formation in Virginia and North Carolina in the eastern United States. The formation consists of cyclical beds of black and grey lacustrine (lake) mudstone and shale. It is a konservat-lagerstätte renowned for its exceptionally preserved insect fossils, along with small reptiles, fish, and plants. Dinosaur tracks have also been reported from the formation.

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Geology

Regional setting

The Cow Branch Formation is exposed in the Dan River-Danville Basin,1213 a narrow half-graben which extends across the border of Virginia and North Carolina in the eastern United States. The basin has also been termed the Danville Basin14 (emphasizing the northern portion in Virginia) or the Dan River Basin15161718 (emphasizing the southern portion, in North Carolina). It is one of many Triassic-Jurassic rift basins stretching from northeast to southwest in eastern North America, collectively described as the Newark Supergroup.19

The Cow Branch Formation was initially distinguished by Meyertons (1963), working in the Virginian portion of the basin. He considered it to be a member of the Leaksville Formation, a name which encompassed almost all Triassic sediment in the basin.20 Thayer (1970), working in North Carolina, split up the Leaksville Formation and raised its members to formation status within the Dan River Group.21 The Cow Branch Formation was divided into upper and lower members, though the lower member has subsequently been renamed to the Walnut Cove Formation.22 Recent revisions place the Cow Branch Formation above the Dry Fork Formation and below the Stoneville Formation.23

The type section of the Cow Branch Formation was a former roadcut along Virginia Route 856, in Pittsylvania County southeast of Cascade.24 A new lectostratotype was proposed in 2015: a large stone quarry extending across the state line by the Dan River near Eden, North Carolina.25 This quarry, commonly known as the Solite Quarry, is technically a cluster of three quarry pits, one in Pittsylvania County, Virginia and two in Rockingham County, North Carolina. The site is home to the most extensive and fossiliferous exposures of the formation. Exceptionally-preserved fossils were first reported from the site in 1978, and collection has continued to the present.262728

Sedimentology and paleoenvironment

The sediments of the Cow Branch Formation are dark grey to black in color and generally fine-grained. Blocky mudstones and thinly-laminated shale are the most common lithologies. The formation is thickest and most fine-grained at the state line, approximately in the middle of its exposed area. Here, the formation is about 1,900 metres (6,200 ft) thick. Coarser sediments such as dark grey sandstone are more prevalent to the southeast and northwest, though periodic black mudstone beds are still frequently encountered. Color is the most useful metric for distinguishing the Cow Branch Formation in the field, as red and purple sediments are practically absent, unlike the Dry Fork and Stoneville formations.29

The Cow Branch Formation represents a lacustrine (lake) system in a warm tropical climate, only around 2°30 to 4°31 north of the equator. Deposition preceded at an estimated rate of around 46.3 cm/kyr.32 Bioturbation is almost completely absent, indicating that the lake bed was uninhabited by burrowing animals. Insect-bearing fossil layers were likely completely freshwater while the fish-bearing layers may have been somewhat saltier. As in modern rift lakes, high water levels could have initiated brine seeps along the edge of the basin, adding sodium into the lake system. Quartz is conspicuously absent even from the siliciclastic layers, having been replaced with albite (high-sodium feldspar) through diagenetic processes.3334

The high frequency of dolomite in the formation indicates that the lake was strongly alkaline, with its water saturated with magnesium supplied from older carbonate rocks in the area.3536 The lack of bioturbation, mudcracks, or root casts has traditionally been taken as evidence that the waters were deep enough to be continually stratified, with the hypolimnion (deepest portion) completely lacking oxygen.373839 An alternative hypothesis suggests that the lake was rather shallow, albeit still deep enough to have been permanent during the formation's deposition. This is supported by the abundance of dolomite, a mineral which forms most easily in salty shallow-water environments. In addition, the insect-bearing layers nearly lack organic carbon, suggesting that the lakebed was fully oxygenated even at its deepest extent.4041 The lake sediments have a high concentration of fluorine, a fact which may help to resolve the near-absence of bioturbators. As with excessive salinity, excessive fluorine can be toxic for fully aquatic organisms (including bioturbators and freshwater plants), but air-breathing insects can persevere and thrive close to the shoreline.4243

Sediment cycles

Sediment cycles are readily apparent in the formation, shifting between the extremes of black microlaminated shale and massive coarse mudstone.44 These are identified as Van Houten cycles, a name applied to fluctuating lake depositional conditions throughout the Triassic rift basins of the Newark Supergroup.45 Each cycle probably corresponds to variations in precipitation tied to the earth's precession, a type of Milankovitch cycle which oscillates on a scale exceeding 21,000 years.46 1747 or 1848 cycles are generally acknowledged in a continuous section at the Solite Quarry. Some sources estimate that up to 30 cycles were preserved at the site, factoring in all three quarry pits combined.49 The semi-precessional (10,000 to 15,000 year) astronomical cycle is another strong influence on sedimentation.50

In the Cow Branch Formation, each cycle begins with a brief package of fine silty claystone.5152 The uppermost portion of this package is rich in mica and carbon, with very little calcareous material. It is followed shortly by a very thin but laterally extensive dolomite bed.5354 These layers correspond to a period of rising lake levels. Fossils of all types are most common in the succeeding black shale and associated lithologies, the point where the lakes are at their deepest extent.555657 Most of the fossiliferous unit is calcareous black shale, though thin beds of extremely fine siliclastic clay can also be found. Insect fossils are predominant in microlaminated carbon-poor shale while fish, plants, and coprolites tend to occur among interbedded carbon-rich dolomite, siltstone, and fine sandstone.5859 The fossiliferous layers occupy only a small portion of each cycle, less than a thousand years' worth of sedimentation.60 They give way to a thicker and coarser series of shallow-water siltstone, first with a high proportion of pyrite and slickensides, then salt casts, and finally massive siltstone beds scoured by wave action.616263

Age

The Cow Branch Formation is certainly from the Late Triassic, though there is disagreement as to the exact age of its sediments. The fossil content and depositional environment are very similar to the Lockatong Formation in the Newark Basin of the Mid Atlantic region, and to a lesser extent the Chinle Formation in the southwest United States.64 During the late 20th century, these formations were often assigned to the later part of the Carnian stage.656667 This was justified by their fossil content, particularly palynomorphs, which were comparable to the Middle Keuper of Germany.6869 The conchostracan Anyuanestheria has been reported from both the German Middle Schilfsandstein and the "lower Cow Branch Formation"70 (now known as the Walnut Cove Formation).71

A Carnian age has been brought into doubt by a more diverse suite of dating methods in the Newark Supergroup.72 Starting in the 1990s, the depositional history of the Newark Basin was recalibrated through a combination of core drilling, radiometric dating, cyclostratigraphy, and magnetostratigraphy. The result was the Newark astrochronostratigraphic polarity time scale (APTS), a unifying system which provides precise ages for sediment layers within the basin.7374 The resulting ages were younger than previously expected. For example, the Lockatong Formation was assigned a mid-Norian age (222.56 – 218.11 Ma), rather than late Carnian.75

The same techniques used to create the Newark APTS can be applied to other basins with continuous cyclical deposition, such as the Dan River-Danville Basin. A magnetostratigraphic sequence has been reconstructed for the Dan River-Danville Basin since 1997, assisting correlation to the Newark Basin.76 There are at least twelve pairs of normal-reverse magnetic polarity chrons recorded in the Dan River-Danville Basin. Four of these magnetostratigraphic intervals were present through the deposition of the Cow Branch Formation: a long reverse chron (D3r), followed by a short normal chron (D4n), a moderate-length reverse chron (D4r) and finally a long normal chron (D5n). These four chrons have been equated with chrons E11r, E12n, E12r, and E13n (respectively) in the Newark Basin.7778 Chrons E11r to E13n apply to a period of time extending from the early-mid Lockatong Formation (Nursery Member) up to the early Passaic Formation (Warford Member), 221.47 Ma to 216.97 Ma.79 If the Cow Branch Formation is equivalent to this interval, then it would be firmly positioned within the Norian stage.80 The fossil beds of the Solite Quarry are in the lower-middle part of the formation, with an estimated age close to 220 Ma.8182

Paleobiota

Color key
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Reptiles

Reptiles of the Cow Branch Formation
Genus / TaxonSpeciesMaterialNotesImages
ApatopusA. sp.Footprints838485Presumed phytosaur footprints
AtreipusA. milfordensis86Footprints87888990Footprints of early quadrupedal dinosaurs or dinosauromorphs, possibly ornithischians.91
GrallatorG. sp.Footprints9293Presumed dinosaur footprints
GwyneddichniumG. sp.Footprints94Footprints, presumably created by small tanystropheids such as Tanytrachelos.95
Mecistotrachelos96M. apeoros97Multiple partial skeletons, two of which have been described98A probable archosauromorph with elongated ribs, presumably hosting gliding membranes.99
RutiodonR. carolinensis100A partial skeleton101 and teeth.102A phytosaurid (or mystriosuchine) phytosaur
Tanytrachelos103T. ahynis104Numerous specimens, including complete skeletons.105A small tanystropheid archosauromorph closely related to Tanystropheus. Some specimens preserve traces of muscles and ligaments near the tail.106107

Fish

Undescribed pholidophorids and holosteans were present.108109 A relatively large freshwater shark tooth is also known from the formation.110

Fish of the Cow Branch Formation
Genus / TaxonSpeciesMaterialNotesImages
DiplurusD. cf. newarki111112At least one large specimen113A coelacanth, sometimes placed in the genus Osteopleurus.114115
CionichthysC. sp.116117A redfieldiiform118
cf. Pariosteguscf. Pariostegus sp.119120121A coelacanth
SemionotusS. brauni.122123124At least three specimens125A semionotiform ginglymodian
SynorichthysS. sp.126127128"Several partial skeletons"129A redfieldiiform130
TurseodusT. spp.131132133At least two specimens134A "palaeoniscid"135136-grade actinopterygian

Arthropods

"Conchostracans" (clam shrimp) from the formation have typically been assigned to the genera Cyzicus and Palaeolimnadia,137138 though they may instead be species of Eustheria.139 Apart from arthropods, other invertebrate fossils from the Cow Branch Formation include (uncommon) Scoyenia burrows and indeterminate unionid bivalves.140

The most abundant insects are aquatic hemipterans (78% of specimens identifiable to order), with beetles in a distant second place (10%).141 Most hemipteran and beetle fossils have yet to be assigned to the species level. In terms of named diversity, dipterans (flies) make up the bulk of the assemblage, despite representing only 1.5% of insect fossils from the Solite Quarry.142 Within the shale layers bearing insect fossils, aquatic insects are most abundant in the early part of a layer, while terrestrial insects maintain a low but steady presence through the entire layer.143

97 different morphotypes (forms) of beetles occur in the Solite Quarry. In terms of size distribution, the beetle fauna is similar to modern ecosystems, with just over half of all morphotypes between 2 and 4 mm in length. Most morphotypes are rare, with 84% known from a single specimen. The two major exceptions are Leehermania and Holcoptera, both of which are known from numerous specimens (about 77% of all beetle fossils from the site). Few morphotypes are shared between Solite and other Triassic-Jurassic insect faunas. Archostematan beetles are rare at Solite despite making up the vast majority of Triassic-Jurassic beetle diversity, as indicated by other sites.144

Arthropods of the Cow Branch Formation
Genus / TaxonSpeciesMaterialNotesImages
Alinka145A. cara146Two specimens.147A procramptonomyiid fly.148
ArchescytinidaeA. indet.149Indeterminate archescytinid thrips.
ArchitipulaA. youngi150"Excellent specimens"151A limoniid crane fly in the subfamily Architipulinae.152
Argyrarachne153A. solitus154A single juvenile specimen missing the abdomen.155An araneomorph spider, one of the oldest known potential species of araneomorph.156
BlattodeaB. indet.157158Indeterminate cockroaches.
Brachyrhyphys159B. distortus160One distorted female specimen.161A protorhyphid fly.162
Cascadelcana163C. virginiana164A wing.165The oldest known member of Elcanidae, a family of cricket-like orthopterans.166
?CrosaphisC. virginiensis167One specimen.168A crosaphidid fly tentatively assigned to Crosaphis.169
cf. Clytiopsiscf. C. sp.170171A crayfish-like decapod.172173
DarwinulaD. spp.174175Freshwater ostracods (seed shrimp).
DipteraD. indet.176177Indeterminate flies, including eoptychopterids and culicomorphs.178
HolcopteraH. solitensis179Two elytra with preserved color patterns.180A water beetle in the family Coptoclavidae.181
Leehermania182L. prorova183Numerous specimens.184A beetle, originally identified as the oldest known staphylinid (rove beetle),185 and later classified as a myxophagan.186
Metarchilimonia187M. krzeminksorum188Two specimens, one of which is a female.189A limoniid crane fly in the subfamily Architipulinae.190
M. solita191One specimen.192A limoniid crane fly in the subfamily Architipulinae.193
MormolucoidesM. articulatusNumerous specimens preserved together in "death beds".194Insect larvae with thick mandibles, likely the aquatic grubs of beetles such as Holcoptera.195
NaucoridaeN. indet.196Indeterminate naucorids (creeping water bugs).
OrthopteraO. indet.197Indeterminate orthopterans (crickets, grasshoppers, and allies)
PhoroschizidaeP. indet.Indeterminate stem-group beetles.
?Phyllocarida?P. indet.198199Possible indeterminate phyllocarid crustaceans.200201 May represent hemipterans or fly pupae instead.202
PhyloblattaP. grimaldii203One of the youngest known phyloblattid cockroaches.204
Prosechamyia205P. dimedia206One specimen.207A stem-group brachyceran fly.208
P. trimedia209One specimen.210A stem-group brachyceran fly.211
Pseudopolycentropodes212P. virginicus213Three specimens.214A scorpionfly in the family Pseudopolycentropodidae.215 Initially misidentified as a trichopteran (caddisfly).216
ThysanopteraT. indet.217Indeterminate thrips.
TipulomorphaT. indet.Indeterminate tipulomorphs (crane flies and allies).
Triassonepa218T. solensis21987 specimens.220The oldest known belostomatid (predaceous water bug).221
Triassopsychoda222T. olseni223One female specimen.224A probable psychodid (drain fly).225
Triassothrips226T. virginicus227Seven specimens.228One of the oldest known thrips.229
VeripleciaV. rugosa230One male specimen.231A paraxymyiid fly.232
Virginiptera233V. certa234Two specimens.235A paraxymyiid fly.236 One specimen was initially misidentified as Crosaphis.237
V. lativentra238One male specimen.239A paraxymyiid fly.240
V. similis241Two male specimens.242A paraxymyiid fly.243
Yalea244Y. argentata245One male specimen.246A procramptonomyiid fly.247
Y. rectimedia248One female specimen.249A procramptonomyiid fly.250

Plants

Plant fossils are abundant. The most common examples are conifer foliage, followed by bennettitales and ginkgophytes. Ferns and sphenophytes are also present, though less common. Two endemic forms, the seed taxon Edenia villisperma and the leaf taxon Pannaulika triassica, have been compared to angiosperms (flowering plants) in their structure, though they likely are unrelated to true angiosperms.251

Plants of the Cow Branch Formation
Genus / TaxonSpeciesNotesImages
BrachyphyllumB. sp.252Abundant conifer leaf-bearing shoots.253
cf. Compsostrobuscf. C. neotericus254255256A conifer seed cone.
CyathoformaC. sp.257Fronds of a large cyatheacean tree fern.258259
cf. Dechellyiacf. D. sp.260261Leaves of an enigmatic gymnosperm, often considered a gnetalean.
DictyophyllumD. sp.262263264Fronds of a dipterid fern.265
Edenia266E. villisperma267A hairy parachuting seed superficially similar to the achenes of modern Platanus (plane trees and American sycamores).268
cf. Elatocladuscf. E. sp.269Conifer leaf-bearing shoots.
EretmophyllumCommon ginkgophyte leaves.270
FraxinopterisF. sp.271Winged gymnosperm seeds.272
GlandulozamitesG. sp.273274Cycad leaves.
cf. Grammaephloioscf. G. sp.275276A lycopod.277
LepacyclotesL. sp.278A lycopod.
"Lepidodendron type"279A lycopod, briefly listed in a single paper on the formation.280
LonchopterisL. virginiensis281282283Fern fronds. The species has sometimes been considered referable to another fern genus, Cynepteris.284
MetreophyllumM. sp.285Leaves of a ginkgophyte similar to Eretmophyllum.286
NeocalamitesN. cf. knowltonii287288289Fragmentary leaf and stem impressions of a sphenophyte (horsetail).
PagiophyllumP. diffusum290Conifer leaf-bearing shoots.
P. simpsoniae291Conifer leaf-bearing shoots.
P. sp.292293294Abundant conifer leaf-bearing shoots.295
Pannaulika296P. triassica297Leaves of an enigmatic plant originally described as similar to dicot angiosperms,298299 though more likely a species of fern.300
PelourdeaP. sp.301Leaves of an enigmatic gymnosperm, possibly a type of conifer.
PodozamitesP. sp.302303304Conifer leaves.
PseudohirmerellaP. delawarensisSeed cones of an early cheirolepid conifer.305306 Fossils of this species from the Solite Quarry were previously known by the names Glyptolepis platysperma307308 or Hirmeriella sp.309
PterophyllumP. cf. Ctenophyllum giganteum.310311Common bennettitale leaves.312
cf. Sagenopteriscf. S. sp.313314Leaves of a caytonialean "seed fern".
SphenobaieraS. sp.315Uncommon ginkgophyte leaves.316
SphenozamitesS. sp.317Bennettitale leaves
ToditesT. gaillardotii?Fronds of an osmundaceous fern. Fossils of this species were initially described as Neuropteris gaillardotii and N. linnaeaefolia.318 The latter species has been reported from the Solite Quarry under the name Acrostichites linnaefolius.319320321 The two species have subsequently been synonymized and referred to the genus Todites.322323
WingateaW. sp.324Fronds of a gleicheniaceous fern.325
cf. Zamiostrobuscf. Z. lissocardus326327328A cycad seed cone.
ZamitesZ. powellii329330331Common bennettitale leaves.332

See also

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Bibliography

References

  1. Olsen, Paul E.; Remington, Charles L.; Cornet, Bruce; Thomson, Keith S. (1978-08-25). "Cyclic Change in Late Triassic Lacustrine Communities". Science. 201 (4357): 729–733. Bibcode:1978Sci...201..729O. doi:10.1126/science.201.4357.729. ISSN 0036-8075. PMID 17750230. S2CID 23168710. https://www.science.org/doi/10.1126/science.201.4357.729

  2. Olsen, Paul E.; Schlische, Roy W.; Gore, Pamela J.W. (1989). Tectonic, Depositional, and paleoecological history of Early Mesozoic rift basins, eastern North America: 28th International Geological Congress Field Trip Guidebook T351 (PDF). Washington D.C.: American Geophysical Union. pp. 1–174. doi:10.1029/FT351. ISBN 0875906583. 0875906583

  3. Liutkus, C. M.; Beard, J. S.; Fraser, N. C.; Ragland, P. C. (2010). "Use of fine-scale stratigraphy and chemostratigraphy to evaluate conditions of deposition and preservation of a Triassic Lagerstätte, south-central Virginia". Journal of Paleolimnology. 44 (2): 645–666. Bibcode:2010JPall..44..645L. doi:10.1007/s10933-010-9445-1. ISSN 0921-2728. S2CID 85464908. http://link.springer.com/10.1007/s10933-010-9445-1

  4. Liutkus-Pierce, Cynthia M.; Fraser, Nicholas C.; Heckert, Andrew B. (2014), Bailey, Christopher M.; Coiner, Lorrie V. (eds.), "Stratigraphy, sedimentology, and paleontology of the Upper Triassic Solite Quarry, North Carolina and Virginia", Elevating Geoscience in the Southeastern United States: New Ideas about Old Terranes—Field Guides for the GSA Southeastern Section Meeting, Blacksburg, Virginia, 2014, Geological Society of America, pp. 255–269, doi:10.1130/2014.0035(09), ISBN 978-0-8137-0035-9, retrieved 2023-12-18 978-0-8137-0035-9

  5. Olsen, Paul E.; Reid, Jeffrey C.; Taylor, Kenneth B.; Whiteside, Jessica H.; Kent, Dennis V. (2015). "Revised stratigraphy of Late Triassic age strata of the Dan River Basin (Virginia and North Carolina, USA) based on drill core and outcrop data". Southeastern Geology. 51 (1): 1–31. doi:10.7916/D82F7MSJ. https://www.researchgate.net/publication/273454856

  6. Olsen, Paul E. (1979). "A new aquatic Eosuchian from the Newark Supergroup (Late Triassic–Early Jurassic) of North Carolina and Virginia" (PDF). Postilla. 176: 1–14. https://www.ldeo.columbia.edu/~polsen/nbcp/olsen_tany_79.pdf

  7. Olsen, Paul E. (1988). "8. Paleoecology and Paleoenvironments of the Continental Early Mesozoic Newark Supergroup of Eastern North America" (PDF). In Manspeizer, Warren (ed.). Triassic-Jurassic Rifting and the Opening of the Atlantic Ocean. Amsterdam: Elsevier. pp. 185–230. doi:10.1016/B978-0-444-42903-2.50013-0. https://www.ldeo.columbia.edu/~polsen/nbcp/olsen_88.pdf

  8. Fraser, Nicholas C.; Grimaldi, David A.; Olsen, Paul E.; Axsmith, Brian (1996). "A Triassic Lagerstätte from eastern North America". Nature. 380 (6575): 615–619. Bibcode:1996Natur.380..615F. doi:10.1038/380615a0. ISSN 0028-0836. S2CID 4360335. https://www.researchgate.net/publication/235735108

  9. Fraser, Nicholas C.; Grimaldi, David A. (1999). "A significant Late Triassic Lagerstaette from Virginia, U.S.A" (PDF). Revista del Museo Civico di Scienze Naturali: Enrico Caffi. 20: 79–84. https://www.museoscienzebergamo.it/wp-content/uploads/2019/01/vol-20s-19.pdf

  10. Fraser, Nicholas C.; Grimaldi, David A. (2003). "Late Triassic continental faunal change: New perspectives on Triassic insect diversity as revealed by a locality in the Danville basin, Virginia, Newark Supergroup". In LeTourneau, Peter M.; Olsen, Paul E. (eds.). The Great Rift Valleys of Pangea in Eastern North America, Volume 2: Sedimentology, Stratigraphy, and Paleontology. New York: Columbia University Press. pp. 192–205.

  11. Weishampel et al., 2004, pp.517-607

  12. Olsen, Paul E.; Schlische, Roy W.; Gore, Pamela J.W. (1989). Tectonic, Depositional, and paleoecological history of Early Mesozoic rift basins, eastern North America: 28th International Geological Congress Field Trip Guidebook T351 (PDF). Washington D.C.: American Geophysical Union. pp. 1–174. doi:10.1029/FT351. ISBN 0875906583. 0875906583

  13. Liutkus-Pierce, Cynthia M.; Fraser, Nicholas C.; Heckert, Andrew B. (2014), Bailey, Christopher M.; Coiner, Lorrie V. (eds.), "Stratigraphy, sedimentology, and paleontology of the Upper Triassic Solite Quarry, North Carolina and Virginia", Elevating Geoscience in the Southeastern United States: New Ideas about Old Terranes—Field Guides for the GSA Southeastern Section Meeting, Blacksburg, Virginia, 2014, Geological Society of America, pp. 255–269, doi:10.1130/2014.0035(09), ISBN 978-0-8137-0035-9, retrieved 2023-12-18 978-0-8137-0035-9

  14. Meyertons, C.T. (1963). "Triassic formations of the Danville basin". Virginia Division of Mineral Resources Report of Investigations. 6: 1–65. https://energy.virginia.gov/commerce/ProductDetails.aspx?productID=2263

  15. Paul A., Thayer (1970). "Stratigraphy and geology of Dan River Triassic basin, North Carolina" (PDF). Southeastern Geology. 12 (1): 1–31. http://libres.uncg.edu/ir/asu/f/SoutheasternGeology_Vol012_No01_1970_M.pdf

  16. Olsen, Paul E.; Remington, Charles L.; Cornet, Bruce; Thomson, Keith S. (1978-08-25). "Cyclic Change in Late Triassic Lacustrine Communities". Science. 201 (4357): 729–733. Bibcode:1978Sci...201..729O. doi:10.1126/science.201.4357.729. ISSN 0036-8075. PMID 17750230. S2CID 23168710. https://www.science.org/doi/10.1126/science.201.4357.729

  17. Fraser, Nicholas C.; Grimaldi, David A. (1999). "A significant Late Triassic Lagerstaette from Virginia, U.S.A" (PDF). Revista del Museo Civico di Scienze Naturali: Enrico Caffi. 20: 79–84. https://www.museoscienzebergamo.it/wp-content/uploads/2019/01/vol-20s-19.pdf

  18. Olsen, Paul E.; Reid, Jeffrey C.; Taylor, Kenneth B.; Whiteside, Jessica H.; Kent, Dennis V. (2015). "Revised stratigraphy of Late Triassic age strata of the Dan River Basin (Virginia and North Carolina, USA) based on drill core and outcrop data". Southeastern Geology. 51 (1): 1–31. doi:10.7916/D82F7MSJ. https://www.researchgate.net/publication/273454856

  19. Olsen, Paul E.; Reid, Jeffrey C.; Taylor, Kenneth B.; Whiteside, Jessica H.; Kent, Dennis V. (2015). "Revised stratigraphy of Late Triassic age strata of the Dan River Basin (Virginia and North Carolina, USA) based on drill core and outcrop data". Southeastern Geology. 51 (1): 1–31. doi:10.7916/D82F7MSJ. https://www.researchgate.net/publication/273454856

  20. Meyertons, C.T. (1963). "Triassic formations of the Danville basin". Virginia Division of Mineral Resources Report of Investigations. 6: 1–65. https://energy.virginia.gov/commerce/ProductDetails.aspx?productID=2263

  21. Paul A., Thayer (1970). "Stratigraphy and geology of Dan River Triassic basin, North Carolina" (PDF). Southeastern Geology. 12 (1): 1–31. http://libres.uncg.edu/ir/asu/f/SoutheasternGeology_Vol012_No01_1970_M.pdf

  22. Olsen, Paul E.; Reid, Jeffrey C.; Taylor, Kenneth B.; Whiteside, Jessica H.; Kent, Dennis V. (2015). "Revised stratigraphy of Late Triassic age strata of the Dan River Basin (Virginia and North Carolina, USA) based on drill core and outcrop data". Southeastern Geology. 51 (1): 1–31. doi:10.7916/D82F7MSJ. https://www.researchgate.net/publication/273454856

  23. Olsen, Paul E.; Reid, Jeffrey C.; Taylor, Kenneth B.; Whiteside, Jessica H.; Kent, Dennis V. (2015). "Revised stratigraphy of Late Triassic age strata of the Dan River Basin (Virginia and North Carolina, USA) based on drill core and outcrop data". Southeastern Geology. 51 (1): 1–31. doi:10.7916/D82F7MSJ. https://www.researchgate.net/publication/273454856

  24. Paul A., Thayer (1970). "Stratigraphy and geology of Dan River Triassic basin, North Carolina" (PDF). Southeastern Geology. 12 (1): 1–31. http://libres.uncg.edu/ir/asu/f/SoutheasternGeology_Vol012_No01_1970_M.pdf

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  26. Olsen, Paul E.; Remington, Charles L.; Cornet, Bruce; Thomson, Keith S. (1978-08-25). "Cyclic Change in Late Triassic Lacustrine Communities". Science. 201 (4357): 729–733. Bibcode:1978Sci...201..729O. doi:10.1126/science.201.4357.729. ISSN 0036-8075. PMID 17750230. S2CID 23168710. https://www.science.org/doi/10.1126/science.201.4357.729

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