Variegation is the appearance of differently coloured zones in the foliage, flowers, and sometimes the stems and fruit of plants, granting a speckled, striped, or patchy appearance. The colors of the patches themselves vary from a slightly lighter shade of the natural coloration to yellow, to white, or other colors entirely such as red and pink. This is caused by varying levels and types of pigment, such as chlorophyll in leaves. Variegation can be caused by genetic mutations affecting pigment production, or by viral infections such as those resulting from mosaic viruses. Many plants are also naturally variegated, such as Goeppertia insignis. Most of these are herbaceous or climbing plants, and are most often species native to tropical rainforests.
Many species which are normally non-variegated are known to display variegation. Their appearance is desirable to enthusiasts, and many such plants are propagated and sold as unique cultivars. However, in individuals where the variegation occurs in normally-photosynthetic cells, the lack of functioning chloroplasts can slow growth rate. Conversely, naturally-variegated plants derive benefits from their appearance, such as improved photosynthetic efficiency in low-light conditions and herbivore deterrence.
The term is also sometimes used to refer to colour zonation in minerals and the integument of animals.